Jô'l
S5Z.7
VAPEX
VOL 15(1)
Trimestriel de la Société Belge de Malacologie
association sans but lucratif
Quarterly of the Belgian Malacological Society
2014
10 MARS
A. Cecalupo & I. Perugia
SOMMAIRE
Articles originaux - Original articles
Cerithiopsidae and Newtoniellidae (Gastropoda: Triphoroidea) from French Polynesia area (South Pacific Océan)
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SOCIETE BELGE DE MALACOLOGIE
A. Cecalupo & I. Perugia
NovAREX 15(1): 1-22, 10 mars 20 14
Cerithiopsidae and Newtoniellidae (Gastropoda: Triphoroidea Gray) from French Polynesia area (South Pacific Océan)
Alberto CECALUPO
Research Associale c/o Acquario e Civica Stazione Idrobiologica di Milano VialeGadio, 2 -20121 Milano (Ml).
- acecalupo(^yahoo.com - - acecalupo@gmail.com -
Ivan PERUGIA
- Via Roncalceci, 152 -48125 Ravenna, loc. Filetto (RA)
- ivanperugia@virgilio.it -
KEYWORDS. Mollusca, Gastropoda, Caenogastropoda, Triphoroidea, French Polynesia.
ABSTRACT. Fifty-seven species of Cerithiopsidae and Newtoniellidae from French Polynesia are recorded and listed, extending their range distribution. A new genus, Aiistralopsis, flfteen new species of Cerithiopsidae and two new species of Newtoniellidae are described.
RESUME. Cinquante-sept espèces de Cerithiopsidae et Newtoniellidae de Polynésie française sont répertoriées et listées, étendant leurs aires de distribution géographique. Un nouveau genre, Amtralopsis, quinze nouvelles espèces de Cerithiopsidae et deux nouvelles espèces de Newtoniellidae sont décrits.
INTRODUCTION
We recently published two books on the Family Cerithiopsidae from the coast of the Central Philippines and Vanuatu identifying 261 new species and establishing 2 new généra (Cecalupo & Perugia, 2012; 2013). The purpose of this paper is to continue the identification of these micro-molluscs from some areas in French Polynesia.
Trôndlé & Boutet (2009) published a list of species from Polynesia reporting only 3 species of Cerithiopsidae and no Newtoniellidae, but they mention 25 additional, unidentified species. This paper allows us to add 74 new records to the list, of which 17 new species and a new genus of Cerithiopsidae.
Number of new species classified by généra
Cerithiopsidae: Amtmlopsis n. gen: 1; Clathropsis: 3; Joculalor: 2; Marshallopsis: 4; Prolixociens: I; Seila: 2; Synthopsis: 2.
Newtoniellidae: Cerithiella: \; Eiimetula: I.
MATERIALS AND METHODS
The material examined was collected during the expéditions conducted by MNHN in the Tropical Indo-Pacific. Specimens were collected during following cruises: Austral Is., N/O “Alis” campagne BENTHAUS 2002, 2011; Austral Is., Rapa, Atelier RAPA (2002); French Polynesia Is. (1990, 1991); Marquesas Is, N/O “Alis” campagne TARASOC ( 1 997); Society Is. N/O “Alis” campagne TARASOC
(2009); Tarava Seamounts Is., Mont Punu Taipu,
N/O “Alis”, campagne TARASOC (2009); Tuamotu Is., N/O “Alis” campagne TARASOC (2009).
We also examined material from the private collections of Jean Letourneux and Michel Boutet (Tahiti) and of Jean Trôndlé (France).
Coll. Michel BOUTET.
Localities; Society Is., Tuamotu Is., number of specimens examined: 37.
Coll. Jean LETOURNEUX.
Localities: Society Is., Tuamotu Is., Marquesas Is., Austral Is., approximately number of specimens examined: 500.
Coll. Jean TRÔNDLÉ.
Localities: Society Is., Tuamotu Is, Marquesas Is., number of specimens examined: 752.
In the generic classification we gave priority to the type of protoconch. The species occasionally lias some different morphological characters than these encountered in the genus in which it is placed. In this case we discuss about this doubtful classification. Only the study of the soft parts and the discovery of additional specimens will clarify these doubts. In this paper we consider appropriate to introduce the new genus Australopsis. Given the known difficulties in the discovery of shells belonging to this family, often withoLit protoconch, and considering that the main purpose of our work is to identify as many species as possible for a géographie area which is not really easily accessible, we decided to introduce new species also based on a single specimen, however, with an intact protoconch and therefore correctiy identifiable.
A. Chcalupü & I. Perugia
Cerithiopsidae and Newtoniellidae from French Polynesia
CLASSIFICATION
Our classification is only based on the shell characters, the soft parts are unknown. We mainly follow the provisional classification of the family proposed by Marshall (1978), using the same distinctive characters.
The size: we use the following terms: very large > 14 mm, large 6^14 mm, medium 3^6 mm, small 2-^3 mm, very small >2 mm. Variation in size is common, particLilarly in Joculator and Horologica.
The protoconch: this is the most important character to identify a species. The protoconch is very fragile, often missing in the available samples, and the lack of its detailed description can make difficult or impossible to identify correctly the species.
The teleoconch: the number of the spiral cords, of the axial ribs and the depth of the suture is important. The shape of the shell, the number of whoiis, the maximum width and the prominence of beads at the intersections of the spiral cords and the axial ribs are also useful. In Horologica species there is occasionally a third spiral cord resulting from the division of the adapical cord.
The colour: it is quite constant and is an excellent character for a preliminary identification of the species; also to consider the range of variability especially for dead collected specimens.
The body whorl: we want to highlight the last whorl, particularly the limit between the base and the columella, a feature that seems to be constant in the specimens belonging to a same genus.
The current classification, updated by Bouchet & Rocroi (2005), differs slightly from that proposed by Marshall (1978).
The superfamily Triphoroidea Gray, 1847 is actually divided into three families: Triphoridae Gray, 1847, Cerithiopsidae H. Adams & A. Adams, 1853 and Newtoniellidae Korobkov, 1955.
The Cerithiopsidae are now divided into 3 subfamilies: Aliptinae Marshall, 1978, Cerithiopsinae H. Adams & A. Adams, 1853 and Seilinae Golikov & Starobogatov, 1975.
The sLibfamily Eumetulinae Golikov & Starobogatov, 1975 is now moved to the family Newtoniellidae.
Abbreviations and text conventions Repositories
MNHN: Muséum national d'Histoire naturelle, Paris, France.
MSNM: Museo Civico di Storia Naturale, Milano, Italy.
LMB: Université Milano Bicocca, Dipartimento Scienze Geologiche e Geotecniche, Milano, Italy.
But: collection Boutet Michel, Papara, Tahiti, Polynésie française.
Ltn: collection Letouneux Jean, Mahina, Tahiti, Polynésie française.
Trn: collection Trôndlé Jean [France], attaché au Muséum national d'Histoire naturelle, Paris.
Other abbreviations
ICZN: International Code of Zoological
Nomenclature.
juv: juvénile form
SEM: Scanning Electron Microscope. spni(s): specimen(s) stn: station
SYSTEMATICS
Family CERITHIOPSIDAE H. Adams & A. Adams, 1853
Genus Australopsis n. gen.
Type species.' Australopsis floresi n. sp.. Austral Is., Rapa.
Diagnosis: shell medium sized, conical, regularly increasing with fiat sides and impressed suture, base moderately constricted. Protoconch of lecithotrophic type, of 2.5 whorls, blunt-tipped; whorls crossed by sub-median granulose carina, fmer grains may extend over the entire surface of whorls.
Remarks. The decision to establish a new genus cornes from the peculiar sculpture of the protoconch that has never been observed in the family Cerithiopsidae; more like the protoconchs of Triphoridae where, however, the généra with dextral shell always hâve a teleoconch with a sculpture of 4 or more spirals. The soft parts are unknown.
Australopsis floresi n. sp.
Figs 1-4, 24-30
Type material. Holotype - MNHN (n° Moll 26460), dim. 5.40 x 1.37 mm, protoconch 0.42 x 0.35 mm, RAPA 2002, stn 34.
Paratype 1 - MNHN (n° Moll 26461), dim. 4.60 x 1.35 mm, protoconch 0.42 x 0.32 mm, RAPA 2002, stn 27.
Paratype 2 - MNHN (n° Moll 26462), dim. 4.65 x
1.26 mm, protoconch 0.43 x 0.35 mm, RAPA 2002, stn 34.
Paratype 3 - MNHN (n° Moll 26462), dim. 4.65 x
1.27 mm, protoconch 0.42 x 0.37 mm, RAPA 2002, stn 34.
Type locality. Austral Is., Rapa, cave at SE Pointe Tematapu, 29 spms, 2-8 m, 27°34,8’S, 144°19,0’W, falling into large cave, muddy bottom, 19 nov. 2002.
Material examined. French Polynesia, Austral Is.:
Rapa, stn 5, south of Tarakoi Is., 1 spm, 8 m, 27°5.6’S, 14.4°18.5’W, dead coral with algae, muddy-
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A. Cecalupo & 1. Perugia
NOVAPEX 15(1); 1-22, 10 mars 20 14
sandy zones, 04 nov. 2002; stn 9, Baie Hiri, 16 spms, 3-24 m, 27°37.3’S, 144°22.2’W, coral, 06 nov. 2002; stn 10, Pointe Komire, 4 spms, 16-18 m, 27°34.8’S, 144°22.8’W, stones eovered with brown algae, 07 nov. 2002; stn 11, N Rapa, Iti Is., 5 spms, 2 m, 27°37.2’S, 144°18.2’W, sandy area on roeky bottom and dead eoral, 07 nov. 2002; stn 16, W Tanna Is., 6 spms, 5 m, 27°36.3’S, 144°18.4”W, mainly dead coral, 09 nov. 2002; stn 17, SW Rarapai Is., 8 spms, 10 m, 27°34.6’S, 144°22.7’W, heap of stones on sandy bottom, 10 nov. 2002; stn 20, Vavai, 4 spms, 5 m, 27°35.4’S, 144°23.3’W, blocks of coral on sandy bottom, 12 nov. 2002; stn 21, E Tapuaki Is., 7 spms, 5 m, 27°34.2’S, 144°20.6’W, blocks of dead coral on sandy bottom, 12 nov. 2002; stn 27, SW Pointe Gotenaonao, 2 spms, 6 m, 27°38.7’S, 144°19.2’W, heap of stones eovered with brown algae, 14 nov. 2002 (1 paratype MNHN); stn 30, Pointe Mei, 2 spms, 16-20 m, 27°38.2’S, 144°18.2’W, dead coral on drop off, 16/18 nov. 2002; stn 32, Vavai, 2 spms, 15-20 m, 27°35.0/35.8’S, 144°22.7/23.0’W, coral, 18/23 nov. 2002; stn 34, cave at SE Pointe Tematapu, 29 spms, 2- 8 m, 27°34,8’S, 144°19,0’W, falling into large cave, muddy bottom, 19 nov. 2002 (holotype MNHN, 2 paratypes MNHN); stn 44, NW Tauna Is., 27 spms, 30 m, 27°36.3’S, 144°18.2’W, drop off with muddy zones, 27 nov. 2002; stn 48, off Pointe Rokuaga, 1 spm, 36 m, 27°34.1’S, 144°22.rW, plateau eovered by sand, 30 nov. 2002.
Description of the holotype. Shell medium in size, conical with convex regularly increasing whorls, slender with fiat sides, last whorl slightly constricted; light-salmon but shell clearly discoloured, base darker. Protoconch of lecithotrophic larval type, blunt-tipped of 2.5 whorls sculptured with one prominent, rounded, sub-median spiral carinae crossed by irregular grains. Teleoconch of 10.5 whorls, perfectly conical, sculpture reticulate of 3 équidistant spiral cords, equal in width, crossed by wider, prosocline axial ribs (about 20 on last whorl). Suture impressed. In last whorl emerging, from insertion of outer lip, fourth spiral cord. Columella short, separated from fiat base of last whorl by furrow; aperture sub-quadrate, outer lip thin and indented by sculpture; siphonal canal open.
Remarks. These Cerithiopsidae from Austral Is. show an interesting protoconch; the many specimens observed show a darker colour with the third spiral cord red brown. Under SEM the surface of the protoconch appears to be minutely granulated throughout. Severns (2011: 236, pl. 101, Figs 5) illustrate an unidentified sample of the genus Melaxia from Hawaii (fam. Triphoridae) with convex whorls and sculpture of 4 spiral cords but with similar protoconch.
Etymology. Dedicated to Heimata Florès, guide and advisor during the worshop "Atelier Rapa".
Genus Clath ropsis Lasaron, 1956 Type species: Clathropsis impedita Laseron, 1956, by original désignation, Masthead Is., Capricorn Group, Qld, Australia,
Clathropsis tuanainaii n. sp.
Figs 6, 33-34
Type material. Holotype - MNHN (n° Moll 26464), dim. 2.32 x 0.75 mm, protoconch 0.46 x 0.27 mm, RAPA 2002, stn 36.
Type locality. Austral Is., Rapa, Pointe Kauria, 1 spm, 27 m, 27°33.5’S, 144°20.8’W, mainly living on coral, 21 nov. 2002.
Material examined. French Polynesia, Austral Is.:
Rapa, stn 36, Pointe Kauria, 1 spm, 27 m, 27°33.5’S, 144°20.8’W, mainly living coral, 21 nov. 2002 (holotype MNHN).
Description of the holotype. Shell juvénile, conical with convex regularly increasing whorls, base exeavate, suture impressed, colour yellowish. Protoconch short, cylindrical, of 2.5 convex whorls sculptured with minute irregular grains stronger in central zone, apex rounded. Colour whitish, semitransparent. Teleoconch of about 6 slightly convex whorls with reticulate sculpture of 3 equal spiral cords crossed by nairower axial ribs (16 on last whorl) forming beads at each intersection with spiral cords. Fourth cord emerging from suture on last whorl. Under SEM limit between columella and base is highlighted by furrow, columella short, crossed by growth Unes throughout. Ridge bordering sub- quadrate aperture with outer lip indented by sculpture. Siphonal canal open.
Remarks. Clathropsis tuanainaii n. sp. is compared with Tubercliopsis cataldinii Cecalupo & Perugia, 2013 (Vanuatu Is.) which has an almost similar protoconch but which can be easily separated by the shape of the teleoconch.
Etymology. Dedicated to Tuanainai Narii, mayor of the village Ahurei which hosted the Rapa workshop team for 6 weeks.
Clathropsis peculiaris n. sp.
Figs 7, 35-36
Type material. Holotype - MNHN (n° Moll 26465), dim. 3.45 x 1.10 mm, protoconch 0.59 x 0.27 mm, TARASOC, stn DW3442.
Type locality. Society Is., between Raiatea & Tahaa, 5 1 5-550 m, 1 6°4 US, 1 5 1 °26’W, 1 6 oct. 2009.
Material examined. French Polynesia, Society Is.:
between Raiatea & Tahaa, N/O “Alis” campagne
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A. ClX'ALUPÜ & 1. PHRUGIA
Cerilhiopsidae and Newtoniellidae irom French Polynesia
TARASOC, stn DW3442, 1 spm, 515-550 m, 16°4rS, 151°26’W, 16oct. 2009 (holotype MNHN)
Description of the holotype. Shell of medium size, conieal with convex, regularly inereasing whorls, tuiTited; base excavate. Protoeonch aeute, eonieal of 4.5 smooth eonvex whorls; last 2.5 with prominent, about SLib-median, broad, rounded carina. Showing under SEM; two apieal whorls granulate, next are smooth exeept for band of minute, irregular granulation on carina; suture marked by thin cord and abapical thin band of small, regularly spaced, grains. ColoLir yellowish, semitransparent. Teleoconch of 6.5 convex whorls with well impressed suture. Reticulate sculpture of 3 spiral cords (first immediately under suture, second thicker), crossed by axial ribs (16 on last whorl). Under SEM: limit between base and columella highlighted by weak groove. Squared tubercles at each intersection of spiral cords with axial ribs. Another narrow beaded spiral at base of last whorl. Columella short, acuminate; aperture with OLiter lip thin, indented by sculpture and wide siphonal canal. White.
Remarks. The protoeonch of Clalhropsis peculiaris has an interesting sculpture while in the diagnosis of the genus (Laseron, 1956) there are no référencés to
sculpture (obviously Laseron could not rely on a SEM analysis). The teleoconch showing ail the characteristics typical of the genus Clalhropsis and, being soft parts unknown, we must consider that the genus présent a range of variability of the sculpture in the protoeonch.
Etymology. Latin, peculiaris, in English peculiar, for the sculpture of its protoeonch.
Clalhropsis payriae n. sp.
Figs 5, 31-32
Type matériel. Holotype - MNHN (n° Moll 26466), dim. 4.40 x 1.25 mm, protoeonch 0.37 x 0.22 mm, RAPA 2002, stn 32.
Paratype 1 - MNHN (n° Moll 26467), dim. 4.20 x 1.12 mm, protoeonch 0.37 x 0.20 mm, RAPA 2002, stn 31. Paratype 2 - MNHN (n° Moll 26468), dim. 4.45 x 1.14 mm, protoeonch 0.37 x 0.21 mm, RAPA 2002, stn 32. Paratype 3 - MNHN (n° Moll 26469), dim. 3.90 x 1.05 mm, protoeonch 0.37 x 0.20 mm, RAPA 2002, stn 44.
Type locality. Austral Is., Rapa, Vavai, 15-20 m, coral, 27°35.0/35.8’S, 144°22.7/23.0’W, 18/23 nov. 2002.
Figures 1-23
1-4. Aiistralopsis floresi n. sp.. Austral Is., Rapa. 1. Holotype MNHN (n° Moll. 26460), 2-8 m [RAPA 2002, stn 34, Cave at SE Tematapu point], 5.40 x 1.37 mm; 2. Paratype 1, MNHN (n° Moll. 26461 ), 6 m [RAPA 2002, stn 27, SW Gotenaonao point], 4.60 x 1.35 mm; 3. Paratype 2, MNHN (n° Moll. 26462), 2-8 m [RAPA 2002, stn 34, Cave at SE Tematapu point], 4.65 x 1.26 mm; 4. Paratype 3, MNHN (n° Moll. 26462), 2-8 m [RAPA 2002, stn 34, Cave at SE Tematapu point], 4.65 x 1.27 mm; 5. Clalhropsis payriae n. sp.. Austral Is., Rapa, holotype MNHN (n° Moll. 26466), 15-20 m [RAPA 2002, stn 32, Vavai], 4.40 x 1.25 mm; 6. Clalhropsis litanainaii n. sp.. Austral Is., Rapa, holotype MNHN (n° Moll. 26464), 15-20 m [RAPA 2002, stn 36, Kauria point], 2.32 x 0.75 mm; 7. Clalhropsis peculiaris n. sp.. Society Is., holotype MNHN (n° Moll. 26465), 15-20 m [TARASOC, stn DW3442], 3.45 x 1.10 mm; 8. Joculalor iUensis n. sp.. Austral Is., holotype MNHN (n° Moll 26472), 2 m [RAPA 2002, stn I I, N Rapa Iti Is.], 1.95 x 0.85 mm; 9. Joculalor bouleli n. sp.. Austral Is., holotype MNHN (n° Moll. 26470), 2 m [RAPA 2002, stn 6, Rapa Is.], 1.67 x 0.77 mm; 10. Marshallopsis bazzocchii n. sp., Marquesas Is., holotype MNHN (n° Moll. 26475), 2 m [MUSORSTOM 9, stn DR1247, Fatu Hiva Is.], 2.02 x 0.82 mm; 11. Marshallopsis Ironcllei n. sp., Tahiti, holotype MNHN (n° Moll. 26479) [stn DR1247, Tahiti Is. Arue], 1.88 x 0.84 mm; 12. Marshallopsis leloumeuxi n. sp., Marquesas Is., holotype MNHN (n° Moll. 26476), 68 m [Ua Pou Haakuti], 1.47 x 0.56 mm; 13. Marshallopsis lahiliensis n. sp., Tahiti Is., holotype MNHN (n° Moll. 26478), 30-60 m [Afaahiti], 1.85 x 0.90 mm; 14. Prolixodens proxima n. sp., Marquesas Is., holotype MNHN (n° Moll. 26480), 340-352 m [MUSORSTOM 9, stn DW 1222, Hiva Oa Is.], 5.30 x 1.50 mm; 15. Seila socielalis n. sp.. Society Is., holotype MNHN (n° Moll. 26484), 720 m, [TARASOC, stn DW3474, Moorea], 3.65 X 1.22 mm; 16. Seila relusa n. sp., Tuamotu Is., holotype MNHN (n° Moll. 26483), 507-607 m [TARASOC, stn DW3373, Kaukura], 3.65 x 1.05 mm; 17. Synlhopsis rapaensis n. sp.. Austral Is., Rapa, holotype MNHN (n° Moll. 26485), 2-8 m [RAPA 2002, stn 34, SE Tematapu], 3.60 x 1.05 mm ; 18 - 19. Synlhopsis vavaiensis n. sp.. Austral Is. Rapa, 18. Holotype MNHN (n° Moll. 26493), 15-20 m [RAPA 2002, stn 32, Vavai], 4.44 x 1.28 mm; 19. Paratype I, MNHN (n° Moll. 26494), 3 m, [RAPA 2002, stn 25, Anatakuri Nako bay], 3.68 x 1.05 mm; 20. Euinelula albachiarae n. sp., Tuamotu Is., Rapa, holotype MNHN (n° Moll. 26491 ), 462-980 m, [TARASOC, stn DW3359, Kaukura], 4.70 x 1 .32 mm; 21 - 23. Cerilhiella francescoi n. sp., Tuamotu, Anaa Is., 21. Holotype MNHN (n° Moll. 26488), 3.45 x 0.90 mm; 22. Paratype 2, MNHN (n° Moll. 26490), Tuamotu, Anaa Is., 3.00 x 0.85 mm; 23. Paratype 1, MNHN (n° Moll. 26489), 507-607 m, [TARASOC, stn DW3373, Kaukura], 2.76 x 0.93 mm.
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A. CF.CALUPO& I. PFRUCilA
Novaphx 15(1): 1-22, 10 mars 20 14
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A. e'i-C'ALUPO & I. PliRUGIA
Cerithiopsiclae and Newtoniellidae from French Polynesia
Material examincd. French Polynesia, Austral Is.:
Rapa, stn 4, Rarapai Is., 3 spms, 1 8 m, heap of stones covered by brown algae, 27°34.3’S, 144°22.rW, 04 nov. 2002; stn 5, south of Tarakoi Is., 1 spm, 8 m, dead corals with algae, muddy sand zones, 27°5.6’S, 14.4°8.5’W, 04 nov. 2002; stn 6, off Baie Aluirei, 13 spms, 42 m, living and dead coral, 27°36.8’S, 144°16.7’W, 05 nov. 2002; stn 8, SE Tauna Is., 5 spms, 52-57 m, roeky bottom with sandy area, 27°36.5’S, 144°17.7’W, 06/22 nov. 2002; stn 9, Baie Hiri, 6 spms, 3-4 m, coral, 27°37.3’S, 144°22.2’W, 06 nov. 2002; stn 31, Pointe Mei, 2 spms, 6 m, heap of stones, 27°38.2’S, 144°18.2’W, 16 nov. 2002 (1 paratype MNHN); stn 32, Vavai, 15 spms, 15-20 m, coral, 27°35.0/35.8’S, 144°22.7/23.0’W, 18/23 nov. 2002 (holotype MNHN, 1 paratype MNHN); stn 44, au NW Tauna Is., 13 spms, 30 m, drop off with muddy zones, falling with muddy past, 27°36.3’S, 144°18.2’W, 27 nov. 2002 (1 paratype MNHN); stn 47, Baie Ahurei bay, 6 spms, 33 m, mud with coral heads, 27°36.7’S, 144°19.rW, 29 nov 2002.
Description of the holotype. Shell medium in size, conical with convex regularly increasing whorls, suture strongly impressed, base excavate. Protoconch near cylindrical, of 3 convex whorls. Under SEM; apex spherical, initially smooth then showing rather regular granulation extended to entire second whori and arranged in 5 lines. Colour yellowish, opaque. Teleoconch of 8.5 convex whorls. Reticulate sculpture of 3 spiral cords crossed by axial ribs (16 on last whori); spirals and ribs equal in size. Beads at each intersection of spiral cords with axial ribs. Fourth spiral at base of last whori. Under SEM: thin close-set axial threads on entire surface; limit between fiat base of spire and columella highlighted by weak groove. Shell covered by thin, transparent periostracum. Columella acuminate, crossed by many growth lines; aperture sub-circular with siphonal canal well detlned. Outer lip thin, indented by sculpture. Background
colour yellowish, semitransparent.
Reniarks. Clalhropsis payriae n. sp. is compared with C. poppeanim and C. zannii, both of Cecalupo & Perugia, 2012 from Central Philippines; the first has the main différence in the protoconch of 4.5 whorls vs 3, C. zannii has a protoconch of 3.5 whorls and an apex with a different, thicker, granulation. It is also compared with C. tuanainaii n. sp. (see below), which has a similar protoconch but shorter, of 2.5 whorls with more marked granulation, and a larger but narrower teleoconch.
Etymology. Named after Claude Payri, Université de Polynésie française, (coordinator of “ Faune et Flore de Rapa”) who very efficiently coordinated this field work in Rapa.
Genus Jociilator Hedley, 1909
Type species: Cerithiopsis ridicula Watson, 1886.
Wednesday Is., Cape York, NE Australia.
Jociilator hoiiteti n. sp.
Figs 9,37-38
Type material. Holotype, MNHN (n° Moll 26470), dim. 1.67 x 0.77 mm, protoconch. 0.22 x 0.19 mm, RAPA 2002, stn 6.
Paratype 1 - MNHN (n° Moll 26471), dim. 1.52 x 0.68 mm, without protoconch. RAPA 2002, stn 6.
Type locality. Austral Is., Rapa, off Ahurei bay, 42 m, 27°36.8’S, 144°16.7’W, living and dead coral, 05 nov. 2002.
Material examined. French Polynesia, Austral Is.:
Rapa, stn 6, off Ahurei bay, 2 spms, 42 m, 27°36.8’S, I44°16.7’W, living and dead coral, 05 nov. 2002 (holotype MNHN, 1 paratype MNHN).
Figures 24 - 34
24 - 30. Australopsis floresi n. sp.. Austral Is., Rapa. 24-25. Holotype MNHN (n° Moll. 26460), 2-8 m [stn 34, Cave at SE Tematapu point], 5.40 x 1.37 mm, protoconch 0.42 x 1.77 mm; 26, 29. Paratype 1, MNHN (n° Moll. 26461 ), 6 m [stn 27, SW Gotenaonao point], 4.60 x 1.35 mm, protoconch 0.42 x 0.32 mm; 27, 30. Paratype 2, MNHN (n° Moll. 26462), 2-8 m [stn 34, Cave at SE Tematapu point], 4.65 x 1 .26 mm, protoconch 0.43 x 0.35 mm; 28. Paratype 3, MNHN (n° Moll. 26462), 2-8 m [stn 34, Cave at SE Tematapu], protoconch 0.42 x 0.37 mm; 31 - 32. Clathropsis payriae n. sp.. Austral Is., Rapa, holotype MNHN (n° Moll. 26466), 15-20 m, [stn 32, Vavai], 4.40 x 1.25 mm, protoconch 0.37 x 0.22 mm.
33 - 34. Clathropsis tuanainaii n. sp.. Austral Is., Rapa, holotype MNHN (n° Moll. 26464), 15-20 m, [stn 36, Kauria point], 2.32 x 0.75 mm, protoconch 0.46 x 0.27 mm.
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Cerilliiopsidae and Newtoniellidae l'rom French Polynesia
Description of the holotype. Shell very small, oval with constricted base; suture not very distinct. Protoconcli very short, near cylindrical of 2.5 smooth convex vvhorls; apex rounded showing, under SEM, very minute granulation; last whorl partially SLibmerged in First whorl of teleoconch. Colour white, opaque. Teleoconch of 4.5 whorls. Reticulate sculpture of 3 spiral cords, about equal in strength, crossed by weaker axial ribs, about 16 on last whorl. Strong beads at each intersection with spiral cords. In last whorl first spiral is slightly stronger and fourth narrower beaded spiral at base. Columella short and broad, obliquely truncate, divided from last whorl with cord, surface crossed by many crisp growth lines. Columellar callus bordering sub-circular aperture; anal SLilcLis and siphonal canal wide and well defined. Colour yellow.
Remarks. Joculator boiiteti n. sp. shows a very short protoconch, typical of the genus Spécula, but its outline is clearly typical of the genus Joculator. It is compared with some similar species, J. mimitus, J. gemmae, J. luteolus, J. perliicidus, J. recisus and J. priorai, ail from Central Philippines (see Cecalupo & Perugia, 2012) with, obviousiy, a main différence in the protoconch. The protoconch of J. bouteti n. sp. is very like to Spécula fragilis Cecalupo & Perugia, 2012 but easily distinguishable by its more conical, turreted teleoconch with a wider suture.
Etymology. Dedicated to Michel Boutet of Tahiti.
Joculator itiensis n. sp.
Figs 8, 39-40
Type material. Holotype - MNHN (n° Moll 26472), dim. 1.95 x 0.85 mm, protoconch 0.25 x 0.18 mm, RAPA 2002 stn 11.
Paratype I - MNHN (n° Moll 26473), dim. 1.78 x 0.78 mm, protoconch 0.22 x 0.20 mm, RAPA 2002, stn 11. Paratype 2 - MNHN (n° Moll 26473), dim. 1 .98 x 0.95 mm, protoconch 0.25 x 0. 1 8 mm, RAPA 2002, stn 11. Paratype 3 - MNHN (n° Moll 26474), dim. 1.55 x 0.76 mm, protoconch 0.23 x 0. 1 8 mm, RAPA 2002, stn 4 1 . Paratype 4 - MNHN (n° Moll 26473), dim. 1.70 x 0.80 mm, protoconch 0.22 x 0. 1 8 mm, RAPA 2002, stn 1 1 . Paratype 5 - MNHN (n° Moll 26473), dim. 1.65 x 0.73 mm, protoconch 0.20 x 0. 1 7 mm, RAPA 2002, stn 11.
Paratype 6 - MNHN (n° Moll 26473), dim. 1.71 x 0.70 mm, protoconch 0.23 x 0.16 mm, RAPA 2002, stn I I.
Type locality. Austral Is., Rapa (Rapa Iti), I spm, 2 m, 27°37.2’S, 144°18.2’W, small sandy zones on fiat rocky bottom et dead coral, 07 nov. 2002.
Material examined. French Polynesia, Austral Is.:
Rapa, stn 4, Rarapai Is., 5 spms, 18 m, 27°34.3’S, 144°22.rW, heap of stones covered with brown algae, 04 nov 2002; stn 6, off Baie Ahurei, 4 spms, 16-20 m, 27°36.8’S, I44°I6.7’W, living and dead coral, 05 nov 2002; stn 1 1, N Rapa Iti Is., 1 spm, 2 m, 27°37.2’S, I44°18.2’W, small sandy zones on fiat rocky bottom et dead coral, 07 nov. 2002 (holotype MNHN, 5 paratypes MNHN); stn 16, W Tauna Is., 6 spms, 5 m, 27°36.3’S, 144°18.4’W, mainly dead coral, 09 nov 2002; stn 25, Baie Anatakuri Nako, 4 spms, 3 m, 27°38.4’S, 144°18.9’W, blocks of dead coral sandy area, 13 nov 2002; stn 28, Pointe Taekateke, 4 spms, 30 m, 27°38.4‘S, 144°20.6’W, lot of stones covered with algae, 15 nov. 2002; stn 30, Pointe Mei, 3 spms, 16-20 m, 27°38.2’S, 144°18.2’W, coral mort on drop off, 16/18 nov. 2002; stn 32, Vavai, 13 spms, 15-20 m, 27°35.0/35.8’S, 144°22.7/23.0’W, coral, 18/23 nov 2002; stn 33, Pointe Teruametitoi, 10 spms, 30 m, 27°34.8’S, 144°I8.6’W, dead coral, 19 nov. 2002; stn 36, Pointe Kauria, 18 spms, 27 m, 27°33.5’S, 144°20,8’W, mainly living coral, 21 nov 2002; stn 41, Baie Anarua, 2 spms, 5 m, 27°36.3’S, I44°22.7’W, coral heads on sandy bottom, 25 nov 2002 ( I paratype MNHN); stn 44, NW Tauna Is., 6 spms, 30 m, 27°36.3’S, 144°18.2’w, drop off with muddy zones, 27 nov. 2002; stn 47, Baie Ahurei, 4 spms, 33 m, 27°36,7’S, 144°19,rW, mud with coral heads, 29 nov. 2002; stn 48, off Pointe Rukuaga, 6 spms, 36 m, 27°34.rS, 144°22.rW, plateau covered by sand, 30 nov. 2002.
Description of the holotype. Shell very small, oval in shape with constricted base, suture moderately impressed. Protoconch short, conical of 3.5 convex whorls, smooth with exception, seen under SEM, for adapical narrow band of microgranules in first 2 whorls; apex rounded. Last whorl partially submerged in first whorl of teleoconch. Colour whitish.
Figures 35 - 44.
35 - 36. Clathropsis peculiari.s n. sp.. Society Is., holotype MNHN (n° Moll. 26465), 15-20 m, [TARASOC, stn DW3442], 3.45 x 1 . 1 0 mm, protoconch 0.59 x 0.27 mm; 37 - 38. Joculator itiensis n. sp.. Austral Is., holotype MNHN (n° Moll. 26472), 2 m, [stn I 1 , N Rapa Iti Is.], 1 .95 x 0.85 mm, protoconch 0.25 x 0. 1 8 mm; 39 - 40. Joculator bouteti n. sp.. Austral Is., holotype MNHN (n° Moll. 26470), 2 m, [stn 6, Rapa Is.], 1 .67 x 0.77 mm, protoconch 0.22 x 0.19 mm; 41 - 42. Marshallopsis bazzocchii n. sp., Marquises Is., holotype MNHN (n° Moll. 26475), 2 in, [stn DR 1247, Fatu Hiva Is.], 2.02 x 0.82 mm, protoconch 0.50 x 0.30 mm; 43 - 44. Marshallopsis letourneiixi n. sp., Marquises Is., holotype MNHN (n° Moll. 26476), 68 m, [Ua Pou Haakuti], 1.47 x 0.56 mm, protoconch 0.50 x 0.30 mm.
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Cerithiopsidae and Newtoniellidae from French Polynesia
Teleoconch of 5 whorls. Sculpture of 3 spiral cords (second narrower) crossed by wider axial ribs, 16 on last whorl. Strong beads at each intersection with spiral cords. Another beaded cord is at base of last whorl. Columella short and broad divided from columella by fourth cord and with surface crossed by many thin crisp folds. Ridge bordering sub-circular aperture. Anal sulcus and wide siphonal canal well defined. Colour red brown, First whorl and columella pale yellowish.
Remarks. Joculator itiensis n. sp. is a shell with a typical outline of Joculator and can be compared with many species from Central Philippines (Cecalupo & Perugia, 2012) as: J. fuscus, J. fiirvus, J. ater, J. carpatinus, J. caliginosus, J. obsciirus, ail with similar colour but different protoconch and J. mimitus, J. pygmaeus, J. perliicidus, J. drivasi, J. recisus, that hâve a different colour. It is also compared with J. lazzarii and J. granulosus Cecalupo & Perugia. 2013 from Vanuatu Is.: both hâve a different teleoconch, the First is more constricted at the base and a larger protoconch, the second has a protoconch of 4.5 whorls and an oval elongated teleoconch.
Etymology. From the type locality.
Genus Marshallopsis Cecalupo & Perugia, 2012 Type species: Marshallopsis albachiarae Cecalupo & Perugia, 2012. Bohol Is., Philippines.
Marshallopsis bazzocchii n. sp.
Figs 10,41-42
Type material. Holotype - MNFIN (n° Moll 26475), dim. 2.02 x 0.82 mm, protoconch 0.50 x 0.30 mm, MUSORSTOM 9, stn DR1247.
Type locality. Marquesas Is., Fatu Hiva, 1150-1250 m, 10°34’S, 138°42’W, 01 nov. 1997.
Material examined. French Polynesia, Marquesas
Is.: Fatu Hiva, N/O “Alis” campagne MUSORSTOM 9, stn DR1247, 1 spm, 1150-1250 m, 10°34’S, 138°42’W, 01 nov. 1997 (holotype MNHN)
Description of the holotype. Shell very small, oval elongated with constricted base. Protoconch conical, acute of 5-5.5 convex whorls with rounded apex. Showing under SEM First 1.5 apical whorls are smooth, next whorls are smooth in upper half where showing narrow band of weak grains under suture while lower half is crossed by thin axial prosocline riblets ( 16-1 8 per whorl). Last whorl partly submerged in First whorl of teleoconch. Colour yellowish, opaque. Teleoconch of 4.5-5 whorls; suture moderately impressed. Reticulate sculpture of 2 about equal spiral cords crossed by narrower axial ribs, 16 on last whorl. Strong bead at each intersection with spiral cords. Fourth beaded cord at base of last whorl. Columella
short and broad with weak cord and surface crossed by thin threads. Clear ridge bordering sub-circular aperture; siphonal canal wide and well defined. Colour orange with First spiral cord darker.
Remarks. Marshallopsis bazzocchii n. sp. is compared with M. atrata Cecalupo & Perugia, 2012 from Central Philippines: same protoconch but teleoconch with 3 spiral cords vs 2 and darker in colour.
Etymology. Dedicated to Piero Bazzocchi member of the “Gruppo Malacologico Romagnolo”, Pesaro, Italy.
Marshallopsis letourneuxi n. sp.
Figs 12, 43-44
Type material. Holotype - MNHN (n° Moll. 26476) dim. 1.47 x 0.56 mm, protoconch 0.50 x 0.30 mm. Paratype 1 - MNHN (n° Moll. 26477) dim. 1.45 x 0.54 mm, protoconch 0.50 x 0.30 mm.
Type locality. Marquesas Is., Ua Pou, Haakuti, 68 m.
Material examined. French Polynesia, Marquesas
Is.: Ua Pou, Haakuti, 68 m, 2 spms, ex coll. Letourneux. (holotype MNHN, 1 paratype MNHN).
Description of the holotype. Shell conical, very small, suture impressed, base slightly constricted. Protoconch conical of 5.5 convex whorls. Under SEM: 1.5 apical whorls smooth, next whorls smooth in upper half and with regularly spaced Fine riblets in lower half, about 18 in last whorl. Suture marked by Fine cord and weak grains. Colour yellowish. Teleoconch conical of 4 whorls with reticulate sculpture of 2 spiral cords (First stronger in last whorl) crossed by narrower axial ribs (about 16 on last whorl). Beads at each intersection with spiral cords. Fourth smooth beaded cord at base of last whorl emerging of insertion of outer lip. Columella short with spiral cord and crossed by many growth lines; clear ridge bordering aperture; siphonal canal well defined. Colour dark orange.
Remarks. This shell could be a juvénile specimen and can be confused with Synthopsis praeacuta Cecalupo & Perugia, 2012 from which it differs mainly by the sculpture of the protoconch.
Etymology. Dedicated to Jean Letourneux of Tahiti.
Marshallopsis tahitiensis n. sp.
Figs 13, 45-46
Type material. Holotype - MNHN (n° Moll. 26478) dim. 1.85 x 0.90 mm, protoconch 0.43 x 0.24 mm.
Type locality. Tahiti, Afaahiti, sédiment, 30-60 m.
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Material examined. French Polynesia, Society Is.:
Tahiti, Afaahiti, sédiment, 30/60 m, 6 spms, protoeonch broken, Jun. 1985 ex coll. Trôndlé. (holotype MNHN).
Description of the holotype. Shell with incomplète aperture, very small, oval, suture moderately impressed, base constricted. Protoeonch eonical of 4- 4.5 convex whorls. Under SEM: 1.5 apical whorls smooth, next whorls smooth in upper half and with regularly spaeed fine prosocline riblets in lower half, about 18 in last whorl. Suture marked by fine cord. Colour yellowish. Teleoconch eonical of 5 whorls with reticulate sculpture of 2 spiral cords (first very stronger and axially elongated in last whorl) crossed by axial ribs (about 18-19 on last whorl). Beads at each intersection with spiral cords. Fourth smooth narrow cord at base of last whorl emerging of insertion of outer lip. Columella short with spiral cord and crossed by many growth fines; clear ridge bordering aperture; outer lip broken. Colour yellowish, with second spiral cord orange.
Remarks. This species is compared with Marshallopsis jolandae Cecalupo & Perugia, 2013 from Vanuatu. The examined sample of M. tahitiensis is smaller in size with an oval feature but the main différence is the colour, having the second spiral cord of teleoconch orange while M jolandae has the first red brown. Other shells with similar colour are Horologica mariani, H. splendida and Jocidator ziliolü ail of Cecalupo & Perugia, 2012 from Central Philippines, easily distinguishable by having a smooth protoeonch.
Etymology. From the type locality.
Marshallopsis troendlei n. sp.
Figs 11,47-48
Type material. Holotype - MNHN (n° Moll. 26479) dim. 1.88 x 0.84 mm, protoeonch 0.39 x 0.25 mm.
Type locality. Tahiti, Arue, Récif du Tombeau du Roi, 22 may 1982.
Material examined. French Polynesia, Society Is.:
Tahiti, Arue, Récif du Tombeau du Roi, I spm, 22 may 1982, ex coll. Trondlé (holotype MNHN).
Description of the holotype. Shell oval, very small, suture moderately impressed, base constricted. Protoeonch eonical of 3.5 convex whorls. Under SEM: apical whorl with an irregular granulation, next whorls smooth in upper 3/4 and with band of regularly spaeed fine riblets in lower part, about 20 in last whorl. Colour yellow'ish. Teleoconch eonical of 5 whorls with reticulate sculpture of2 spiral cords (first stronger and, in last whorl, divided by furrow) crossed by narrower axial ribs (about 14 on last whorl). Beads
at each intersection with spiral cords. Fourth smooth beaded cord at base of last whorl emerging of insertion of outer lip. Columella short with clear spiral cord and crossed by many growth fines; ridge bordering aperture; siphonal canal and anal sulcus well defined. Colour pale brown with first 2 whorls and columella yellowish.
Remarks. Similar in colour to Jocidator alligatus Cecalupo & Perugia, 2012 from the Philippines and J. i tiens is, here reported from French Polynesia, but it is smaller then J. alligatus and has a completely different protoeonch from both species.
Etymology. Dedicated to Jean Trôndlé of Fa Force (France).
Genus Prolixodem Marshall B. A., 1978
Type species: Cerithiopsis infracolor Faseron, 1951.
Off Fong Reef, NSW, Australia.
Prolixodens proxima n. sp.
Figs 14, 49-50
Type material. Holotype - MNHN (n° Moll 26480), dim. 5.30 x 1.50 mm, protoeonch 0.62 x 0.27 mm, MUSORSTOM 9, stn DW 1222.
Paratype 1 - MNHN (n° Moll 26481), dim. 2.25 x 0.72 mm, protoeonch 0.62 x 0.27 mm, MUSORSTOM 9, stn DR 1200.
Paratype 2 - MNHN (n° Moll 26482), dim. 4.10 x 1 .25 mm, protoeonch 0.62 x 0.26 mm. Atelier MARQUISES 1997, stn 24 bis.
Type locality. Marquesas Is., Hiva Oa, 340-352 m, 9M4’S, 138°51’W, 30, aug. 1997.
Material examined. French Polynesia, Marquesas
Is.: Hiva Oa, N/O “Alis” campagne MUSORSTOM 9, stn DR 1200 1 spm, 340-352 m, 9°44’S, 138°5rW, 30 aug. 1997 (holotype MNHN); stn DR1200, 1 spm, 96- 100 m, 9°49.9’S, 139°08.9’W, 28 aug. 1997; Atelier MARQUISES 1997 (1 paratype MNHN); stn 24 bis, NW coast of Baie Haahue, 1 spm, 20-34 m, 8°53.60’S, 139°37.00’W, sandy bottom, 10 oct. 1997 ( 1 paratype MNHN).
Description of the holotype. Shell of medium size, eonical with convex regularly increasing whorls, turreted, base excavate. Protoeonch acute, eonical of 4.5 smooth, convex whorls. Under SEM: 1.5 apical whorls granulate, suture defined by thin cord with 2 séries of small grains, those adapical more minute. Next whorls are smooth in upper one-third and with regularly spaeed fine prosocline riblets in lower two- thirds. Colour whitish, semitransparent. Teleoconch of 9 convex whorls with well impressed suture. Reticulate sculpture of 3 main equal spiral cords (first immediately under suture) crossed by axial ribs of same strength (about 22 on last whorl). Squared
A. Cfcalupo & 1. Pprugia
Cerithiopsidae and Newtoniellidae from French Polynesia
tubercles at each intersection with spiral cords. Under SEM: limit betvveen base and columella highiighted by weak groove. Another narrow spiral cord at base of last whorl. Columella short, acuminate; outer lip thin, indented by sculpture, siphonal canal open, well defined. Colour whitish, suture and First spiral yellowish.
Remarks. For the sculpture of the protoconch it is compared with Prolixodens captiosa Cecalupo & Perugia, 2012 from Central Philippines. P. proxima n. sp. has a shorter protoconch, a teleoconch with a less open suture; the shell is also less slender, reaching a larger diameter in width. In those two species the protoconch has a different sculpture from the diagnosis of the genus Prolixodens (Marshall, 1978) "Subséquent whorls with granules on shoulder and fine, crisp, regularly spaced riblets on prosocline sides." (add riblets from suture to suture). Even the base "Base fairly evenly contracted at maturity, smooth." looks different and does not présent any contraction in our species. We consider provisional the assignment to the genus Prolixodens.
Etymology. Latin proxima, for its resemblance with ^Prolixodens captiosa.
Genus Sella A. Adams, 1861
Type species: Triphoris dextroversns A. Adams & Reeve, 1850. China seas.
Sella refusa n. sp.
Figs 16, 53-54
Type material, Flolotype - MNFIN (n° Moll 26483), dim. 3.65 x 1.05 mm, protoconch 0.45 x 0.50 mm, TARASOC, stn DW3373. Dead sample.
Type locality, Tuamotu Is, Kaukura, 507-607 m, 15°41’S, 148°54’W, 04 oct. 2009.
Material examined. French Polynesia, Tuamotu
Is.: Kaukura, N/O “Alis” campagne TARASOC, stn DW3373, 1 spm, 507-607 m, 15°4rS, 148°54’W, 04 oct. 2009 (holotype MNHN).
Description of the holotype. Shell medium in size, conical with Hat sides, suture not easily discernible, colour opaque white. Protoconch of lecithotrophic larval type, blunt-tipped, merging insensibly into teleoconch but of about 2.5 whorls, conical with same inclination of teleoconch and sculptured with one prominent médian smooth spiral carinae; dim. about 0.43 X 0.50 mm. Teleoconch of 7 whorls, sculptured with three prominent smooth spiral carinae, second very narrower. Showing under SEM at base of last whorl, two other thin spiral emerging from suture; in intervals between carinae very thin, close-set axial lamellae; limit between base of shell and columella highiighted by groove. Columella short, obliquely truncated; aperture sub-circular with siphonal canal wide.
Remarks. Few species can be compared with Seila refusa n. sp.: Seila (Lyroseila) cincta (Hutton, 1885) from N. Zealand, a species living between 6 and 30 m depth, illustrated by Marshall (1978), has a flatter protoconch, a more conical teleoconch and a different colour (reddish to yellowish brown). Hutton (1885) included S. cincta in the genus Bittium. Seila (Lyroseila) dilecta Marshall, 1978, from New Zealand, living in a greater depth, up to 805 m, has a similar outline to S. refusa n. sp. but a different colour (yellowish brown, spiral cords white) and a more rounded, not blunt-tipped protoconch.
Etymology. Latin refusa, for the apical truncated form.
Sella socletatls n. sp.
Figs 15, 51-52
Type material. Holotype - MNHN (n° Moll 26484), dim. 3.65 x 1.22 mm, protoconch 0.47 x 0.40 mm, TARASOC, stn DW3374. Dead sample.
Type locality. Society Is., Moorea, 720 m, 17°28’S, 151°26’W,2I oct. 2009.
Figures 45 - 54.
45 - 46. Marshallopsis tahitiensis n. sp., Tahiti Is., holotype MNHN (n° Moll. 26478), 30-60 m, [Ataahiti], 1.85 X 0.90 mm, protoconch 0.43 x 0.24 mm; 47 - 48. Marshallopsis frondlei n. sp., Tahiti, holotype MNHN (n° Moll. 26479), [stn DRI247, Tahiti Is. Arue], 1.88 x 0.84 mm, protoconch 0.39 x 0.25 mm; 49 - 50. Prolixodens proxima n. sp., Marquises Is., holotype MNHN (n° Moll. 26480), 340-352 m, [Hiva Oa Is.], 5.30 x 1.50 mm, protoconch 0.62 x 0.27 mm; 51-52. Seila societatis n. sp.. Society Is., holotype MNHN (n° Moll. 26484), 720 m, [TARASOC, stn DW3474, Moorea], 3.65 x 1 .22 mm, protoconch 0.47 x 0.40 mm; 53 - 54. Seila refusa n. sp., Tuamotu Is., holotype MNHN (n° Moll. 26483), 507-607 m, [TARASOC, stn DW3373, Kaukura], 3.65 x 1 .05 mm, protoconch 0.45 x 0.50 mm.
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Cerithiopsidae and Newtoniellidae l'rom French Polynesia
IVlaterial exaniined. French Polynesia, Society Is.:
Moorea, N/O “Alis” campagne TARASOC, stn DW3474, 1 spm, 720 m, ]7°28’S, 15r26’W, 21 oct. 2009 (holotype MNHN).
Description of the holotype. Shell medium in size, conical with slightly constricted base, suture indistinct, colour opaque white, probably yellowish in tresh samples. Protoconch short of 2.5 smooth, convex whorls. Teleoconch of 7 whorls with spiral sculpture of 3 smooth spiral cords (adapical cord stronger, second very narrower). Under SEM: fine very dose- set axial lamellae in their intervals, well distinct. Limit between base of shell and columella highlighted by groove. At base of last whorl other two thin spirals emerging from suture. Columella short and broad, obliquely truncate. Aperture and outer lip damaged.
Remarks. SeÜa societatis n. sp. is compared with Seila (Lyroseila) dilecta Marshall, 1978 and Seüa (Lyroseila) cincta (Hutton, 1885) both from New Zealand; it is easily distinguished by having a smooth protoconch without keels.
Etymology. From the type locality.
Genus Synthopsis Laseron, 1956
Type species: Synthopsis cylindrica Laseron, 1956.
Michaelmas Cay, Qld, Australia.
Synthopsis rapaensis n. sp Figs 17, 55-56
Type material. Holotype - MNHN (n° Moll 26485), dim. 3.60 x 1.05 mm, protoconch 0.50 x 0.26 mm, RAPA 2002, stn 34.
Paratype 1 - MNHN (n° Moll 26486), dim. 3.55 x 1.00 mm, protoconch 0.50 x 0.21 mm, RAPA 2002, stn 44. Paratype 2 - MNHN (n° Moll 26486), dim. 3.70 x 1.15 mm, protoconch 0.50 x 0.22 mm, RAPA 2002, stn 44. Paratype 3 - MNHN (n° Moll 26487), dim. 4.00 x 1.05 mm, protoconch 0.50 x 0.23 mm, RAPA 2002, stn 44.
Type locality. Austral Is., Rapa, cave SE Pointe Tematapu, 26 spms, 2-8 m, 27°34,8’S, 144°19.0’W, falling inside large cave, muddy bottom, 19 nov. 2002.
Material examined. French Polynesia, Austral Is.:
Rapa, stn 4, Rarapai Is., 1 spm, 18 m, 27°34.3’S, 144°22.rW, blocks of coral covered with brown algae, 04 nov. 2002; stn 8, S/E Tauna Is, 5 spms, depth 52-57 m, 27°36.5’S, 144°17.7’W, on sandy area, 06 nov. 2002; stn 16, W Tauna Is., 1 spm, 5 m, 27°36.3’S, 144°18.4’W, manly dead coral, 09 nov. 2002; stn 21, E Baie Tupuaki, 4 spm, 5 m, 27°34.2’S, 144°20.6’W, blocks of dead coral on sandy bottom, 12 nov. 2002; stn 25, Baie Anatakuri Nako, 6 spms, 3 m, 27°38.4’S, 144°18.9’W, blocks of dead coral on sandy zones, 13 nov. 2002; stn 32, Vavai, 1 spm, 15-20 m, 27°35.0/35.8’S, 144°22.7/23.0’W, coral, 18/23 nov. 2002; stn 34, cave SE Pointe Tematapu, 26 spms, 2-8 m, 27°34,8’S, 144°19.0’W, falling inside large cave, muddy bottom, 19 nov. 2002 (holotype MNHN); stn 38, N Baie Anatakuri, 1 spm, 2 m, 27°37.4’S, 144°18.4’W, sédiment on big rock, 22 nov. 2002; stn 44, NW Tauna Is., 27 spms, 30 m, 27°36.3’S, 144°18.2’W, drop off with muddy zones, 27 nov. 2002 (3 paratypes MNHN); stn 47, Baie Ahurei, 5 spms, 33 m, 27°36,7’S, 144°19,rW, mud with coral heads, 29 nov. 2002; stn 67, Baie Pariati, 1 spm, 3-4 m, 27°34.7’S, 144°21.7’W, sand, mud and algae, 18 nov. 2002.
Description of the holotype. Shell medium in size, nearly cylindrical with slightly constricted base. Protoconch conical of 3.5 convex whorls. Under SEM: apex rounded, initially smooth then with fine granulation extended to lower part of second whorl, roughly arranged in 6-7 Unes; suture marked by minute grains. Colour opaque white, suture orange. Teleoconch of 8 whorls, suture moderately impressed. Reticulate sculpture of 3 spiral cords (second lightly narrower) crossed by weaker axial ribs, 18-19 on last whorl, adapical spiral cord larger. Strong beads at each intersection with axial ribs. Another beaded cord at base of last whorl. Under SEM: intervals between spirals crossed by thin close-set axial threads; limit between base of shell and columella is highlighted by clear groove. Columella short and broad, obliquely truncate, with surface crossed by growth Unes throughoLit. Columellar callus bordering sub-circular aperture with wide siphonal canal well defined. Ground colour orange with first spiral cord white.
Figures 55 - 65
55 - 56. Synthopsis mpaensis n. sp.. Austral Is., Rapa, holotype MNHN (n° Moll. 26485), 2-8 m, [stn 34, SE Tematapu], 3.60 x 1 .05 mm, protoconch 0.50 x 0.26 mm; 57 - 58. Synthopsis vavaiensis n. sp.. Austral Is. Rapa, holotype MNHN (n° Moll. 26493), 15-20 m, [stn 32, Vavai], 4.44 x 1.28 mm, protoconch 0.52 x 0.25 mm;
59 - 60. Eumetula cdhachiarcw n. sp., Tuamotu Is., Rapa, holotype MNHN (n° Moll. 26491), 462-980 m, [TARASOC, stn DW3359, Kaukura], 4.70 x 1.32 mm, protoconch 0.37 x 0.45 mm; 61 - 65. Cerithiella francescoi n. sp., Tuamotu, Anaa Is. 61. Holotype MNHN (n° Moll. 26488), 3.45 x 0.90 mm, 62. Protoconch holotype 0.60 x 0.23 mm; 63. Protoconch paratype 2, 0.60 x 0.22 mm; 64. Protoconch Paratype I, 0.60 x 0.27 mm; 65. Paratype I, MNHN (n° Moll. 26489), 507-607 m, [TARASOC, stn DW3373, Kaukura], 2.76 x 0.93 mm, protoconch 0.60 x 0.23 mm.
14
A. Cecalupo & I. Perugia
Novapex 15(1): 1-22, 10 mars 20 14
15
A. ClX’ALUPO & 1. PERUGIA
Cerithiopsidae and Newtoniellidae from French Polynesia
Remarks. Synfhopsis rapaensis n. sp. is compared with S. enzae Cecalupo & Perugia, 2012 from Central Philippines. It is similar in size and eolour, with the First spiral eords of the teleoeonch white, but it differs in having a protoeonch of 3.5 whorls, with seulpture of minute grains in the first two whorls, vs 4.5 elongated, smooth whorls in S. enzae.
Etymology. From the type loeality.
Synthopsis vavaiensis n. sp.
Figs 18-19, 57-58
Type material. Holotype - MNHN (n° Moll. 26493) dim. 4.55 x 1.28 mm, protoeonch 0.52 x 0.25 mm, RAPA 2002, stn 32.
Paratype 1 - MNFIN (n° Moll. 26494) dim. 3.68 x 1.05 mm, protoeonch 0.52 x 0.25 mm, RAPA 2002, stn 25. Paratype 2 - MNHN (n° Moll 26495), dim. 3.60 x 1.00 mm, protoeonch 0.50 x 0.25 mm, RAPA 2002, stn 44. Paratype 3 - MNHN (n° Moll 26496), dim. 2.90 x 0.90 mm, protoeonch 0.52 x 0.25 mm, RAPA 2002, stn 8.
Type loeality. Austral Is., Rapa, Vavai, 15-20 m, 27°35.0/35.8’S, 144°22.7/23.0’W, coral, 18/23 nov. 2002.
Material examinée!. French Polynesia, Austral Is.:
Rapa, stn 5, south of Tarakoi Is., 1 spm, 8 m, 27°5.6’S, 14.4°18.5’W, dead coral with algae, muddy- sandy zones, 04 nov. 2002; stn 8, SE Tauna Is., 9 spms, 52-57 m, 27°36.5’S, 144°17.7’W, rocky bottom with sandy area, 06/22 nov. 2002 ( 1 paratype MNHN); stn 9, Baie Hiri, 2 spms, 3-24 m, 27°37.3’S, 144°22.2’W, coral, 06 nov. 2002; stn 10, Pointe
Komire (+ 79 m), 1 spm, 16-18 m, 27°34.8’S,
144°22.8’W, stones covered with brown algae, 07 nov. 2002; stn 16, W Tauna Is., 1 spm, 5 m,
27°36.3’S, 144°18.4’W, mainly dead coral, 09 nov. 2002; stn 20, Vavai, 3 spms, 5 m, 27°35.4’S,
144°23.3’W, blocks of coral on sandy bottom, 12 nov. 2002; stn 25, Baie Anatakuri Nako, 1 spm, 3 m, 27°38.4’S, 144°18.9’W, blocks of dead coral on sandy area, 13 nov. 2002 (1 paratype MNHN); stn 32, Vavai, 2 spms, 15-20 m, 27°35.0/35.8’S, 144°22.7/23.0’W, coral, 18/23 nov. 2002 (holotype MNHN); stn 43, Baie Ahurei, I spm, 45 m, 27°36.8’S, 14.4°18.3’W, drop off with muddy bottom at the base, 26 nov. 2002; stn 34, cave SE Pointe Tematapu, 1 spm, 2-8 m, 27°34,8’S, 144°19.0’W, falling inside large cave, muddy bottom, 19 nov. 2002; stn 44, NW Tauna Is., 6 spms, 30 m, 27°36.3’S, 144°I8.2’W, drop otf with muddy zones, 27 nov. 2002
(1 paratype MNHN); stn 47, Baie Ahurei, I spm, 33 m, 27°36.7’S, I44°l9.rw, mud bottom with coral heads, 29 nov. 2002; stn 48, off Pointe Rukuaga, 1 spm, 36 m, 27°34.I’S, I44°22.I’W, plateau covered by sand, 30 nov. 2002.
Description of the holotype. Shell medium in size.
perfectly conical with fiat sides and constricted base. Ground eolour orange. Protoeonch siender, conical of 3.5 smooth convex whorls with same inclination of teleoeonch. Apex spherical, seen under SEM with slight granulation. Teleoeonch of 9.5 fiat whorls. Suture moderately impressed. Reticulate sculpture of 3 spiral cords crossed by axial ribs, 18-19 on last whorl. Spirals and ribs of about equal strength; adapical spiral cord stronger on last whorl. Under SEM: thin close-set threads in intervals of spirals; limit between base of shell and columella highlighted by clear groove. Beads at each intersection of spiral cords with axial ribs. Another cord at base of last whorl emerging from suture. Columella short and broad, obliquely truncate, with surface crossed by many growth lines. Columellar callus bordering sub- circular aperture with wide and well defined siphonal canal.
Remarks. Synthopsis vavaiensis n. sp. is compared with Cehthiopsis powelli Marshall, 1978 from New Zealand which has a similar conical shape. The main différence is localized in the protoeonch; C powelli shows a paler protoeonch of 4.5-5 whorls with a sculpture from minute granules, on the first 1 .5 embryonic whorls, to broken prosocline riblets on the last. In S. vavaiensis n. sp. the protoeonch is of 3.5 smooth whorls with only the embryonic whorls slightly granulate and the sutures marked by a sériés of minute grains.
Etymology. From the type loeality.
Family NEWTONIELLIDAE Korobkov, 1955 Genus Cerithiella Verrill, 1882
Type species: Cerifhium metula Lovén, 1846. Bergen, Norway.
Cerithiella francescoi n. sp.
Figs 21-23, 61-65
Type material. Flolotype - MNHN (n° Moll 26488), dim. 3.45 x 0.90 mm, protoeonch 0.60 x 0.23 mm, Tuamotu Is., Anaa.
Paratype 1 - MNHN (n° Moll 26489), dim. 2.76 x 0.93 mm, protoeonch 0.60 x 0.27 mm, Tuamotu Is., Kaukura, TARASOC, stn DW3373.
Paratype 2 - MNHN (n° Moll 26490), dim. 3.00 x 0.85 mm, protoeonch 0.60 x 0.22 mm, Tuamotu Is., Anaa.
Type loeality. Tuamotu Is., Anaa, sédiment on the external beach.
Material examined. French Polynesia, Tuamotu
Is.: Anaa, sédiment on the external beach, 40 spms, coll. Trôndlé (holotype MNHN, 1 paratype MNHN); stn DW3373, Tuamotu Is., Kaukura, N/O “Alis”, campagne TARASOC, 1 spm, 507-607 m, 15°41’S, I48°54’W, 04 oct2009 (1 paratype MNHN).
Description of the holotype. Shell medium in size.
16
A. CHCALUPO & 1. PliRUGIA
NovAi^EX 15(1); I -22, 1 0 mars 20 1 4
regularly conical witli distinct suture, last whori slightly constricted, light white in colour. Protoconch cylindrical oi' 2.5 smooth convex whorls, apex bulbous. Teleoconch or9.75 convex whorls. Reticular sculpture of 3 spiral eords (adapical eord narrower) crossed by weaker axial ribs (about 16 on last whori); strong roLinded beads on each intersection. Distance between first 2 cords very small; Iburth spiral cord emerging from outer lip. Columella short, separated l'rom base of teleoconch by llllh smooth cord. Columellar callus bordering circulai' aperture; siphonal canal well defmed, anal sulcus barely hinted, margin of outer lip little eorroded.
Remarks. It is eompared with Cerit/iiella cucuminacea (Hedley & Petterd, 1906) from Australia. The drawing of C. cucuminacea shows a more bulbous apex of 1.5 whorls, the suture of the teleoconch is indistinct and there is an obvions columellar fold, details that differentiate it from Cerithiel/a fcancescoi n. sp.
Etymology. Dedicated to Francesco, grandson of Alberto Cecalupo.
Genus Eiimetula Thiele, 1912
Type species: Eumetu/a dilecta Thiele, 1912, by
monotypy. Antarctic.
Emuetula alhachiarae n. sp.
Figs 20, 59-60
Type material. Holotype - MNFIN (n° Moll 26491), dim. 4.70 x 1.32 mm, protoconch 0.37 x 0.45 mm, TARASOC, stn DW3359.
Paratype 1 - MNHN (n° Moll 26492), dim. 4.45 x 1.25 mm, protoconch 0.37 x 0.45 mm, TARASOC, stn DW3373.
Paratype 2 - MNHN (n° Moll 26492), dim. 7.50 x 1 .62 mm, protoconch 0.37 x 0.45 mm, TARASOC, stn DW3373.
Type locality. Tuamotu Is., Kaukura, 462-980 m, 15°57’S, I47°08’W, 01 oct. 2009.
Mateial examinée!. French Polynesia, Tuamotu Is.:
Kaukura, N/O “Alis” campagne TARASOC. stn DW3359, 1 spm, 462-980 m, 15°57’S, I47°08’W, 01 oct. 2009 (holotype MNHN); stn DW3373, 2 spms, 507-607 m, I5°4rs, 146°54’W, 04 oct. 2009 (2 paratypes MNHN).
Description of the holotype. Shell slightly conical with convex regularly increasing whorls, distinct suture, vitreoLis, semitransparent, colourless. Protoconch very short of about 1.5 smooth whorls, not demarked from spire. Teleoconch of 7 convex whorls, smooth except for many fine growth Unes, sculptured with 10 axial prominent ribs from suture to suture, rather orthocline. Aperture sub-circular, outer lip thin.
columella acuminate, smooth.
Remarks. No comparable species from our bibliographical research.
Etymology. Dedicated to Albachiara, grand-daughter of Ivan Perugia.
OTHER MATERIAL EXAMINED FROM PRIVATE COLLECTIONS
These records extend the range distribution Irom the type locality up to French Polynesia.
Clathropsis multispirae Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Society Is. - Tahiti, Mahina, Baie de Matavai, 4 spms, 5 m, 01/1980 [Trn]; Tahiti, Mahina, Baie de Matavai, 20 m [Ltn]; Tahiti, Mahina, reef sédiment, 3 spms, 06/1980 [Trn]; Tahiti, Mahina, Orofara, sédiment on the beach, 1 spm [But]; Tahiti, Papenoo, 1 m [Ltn]; Moorea [Ltn]; Tahiti, Papara, rock brushing, 1 spm, 0.50 m, 30/11/89 [Trn]; Tetiaroa [Ltn]; Mehetia, 55 m, 20 spms [Ltn]; Tuamotu Is. - Rangiroa, Otepipi, sédiment on external beach, 2 spms, 08/2001 [Trn]; Makemo, Passe Arikitamiro, sand under dead coral block, 1 spm, 1 m [Ltn].
Marquesas Is. - Ua Pou, Haakuti, 68 m, 4 spms; Ua Pou, Hakahetau 5 m [Ltn].
Clathropsis poppearum Cecalupo & Perugia, 2012
Type locality: Mactan Is., Philippines.
French Polynesia: Society Is. - Tahiti, Arue, chanel, lagoon, 12 m [Ltn]; Tahiti, Arue, outer slope, 35 m [Ltn]; Tahiti, Mahina, Pointe Vénus, Banc du Dauphin, 9 spms, 18 m [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn].
Clathropsis pulchella Cecalupo & Perugia, 2012
Type locality: Bohol Is., Philippines.
French Polynesia: Society Is. - Tahiti, Mahina, Baie de Matavai, 20 m [Ltn]; Tahiti, Papenoo, 1 m [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia. 55 m, 20 spms [Ltn].
Clathropsis zannii Cecalupo & Perugia, 2012
Type locality: Mactan Is., Philippines.
French Polynesia: Society Is. - Tahiti, Arue, outer slope, 2 spnis, 20-57 m [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia, [Ltn].
17
A. ClX'AL.liPO& 1. Perugia
Cerithiopsidae and Newtoniellidae from French Polynesia
Horologica acuta Cecalupo & Perugia, 2013
Type locality: Espiritu Santo Is., Vanuatu.
French Polynesia: Tuamotu Is. - Makemo, Pouheva reet, Arikitamiro, 1 m, sédiment between dead eoral blocks, 1 spm [Ltn].
Horologica alligata Ceealupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Tuamotu Is. - Rangiroa, near Passe de Tiputa, 1 spm, 100 m [Ltn].
Horologica diffusa Cecalupo & Perugia, 2012
Type locality: Cebu Is., Philippines.
French Polynesia: Society Is. - Tahiti, Tiarei, sédiment, 1 spm, 1 m [Ltn]; Moorea[Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn].
Horologica flava Cecalupo & Perugia, 2013
Type locality: Espiritu Santo Is., Vanuatu.
French Polynesia: Society Is. - Bora Bora, sédiment from outer slope of the reef barrier, 2 spms [But].
Horologica fraudulenta Cecalupo & Perugia, 2013
Type locality: Espiritu Santo Is., Vanuatu.
French Polynesia: Society Is. - Tahiti, Faaone, reef along the coast, 28 spms, 23/05/85 [Trn]; Tahiti, Mitirapa, sédiment on the beach, 2 spms [But]; Tahiti, Passe de Papeete, reef sédiment, 20 spms, 1980/1981 [Ltn]; Tahiti, Punaauia, reef, 16 spms, 1980/81 [Ltn]; Tahiti, Mahina, Orofora, sédiment on the beach, 3 spms, 01/2009 [Ltn]; Moorea [Ltn]; Mehetia [Ltn]; Tetiaroa, beach sédiment [Ltn]; Bora Bora reef, 2 spms, 1 m [Ltn]; Tuamotu Is. - Hao, sédiment on the external beach, 1 spm, 1981 [Ltn].
Horologica infuscata Cecalupo & Perugia, 2012
Type locality: Mactan Is., Philippines.
French Polynesia: Society Is. - Tahiti, Papara, lagoon, sandy bowls near reef, I spm [But]; Tahiti, Mitirapa, sédiment on the beach, 2 spms [But],
Horologica iucunda Cecalupo & Perugia, 2013
Type locality: Espiritu Santo Is., Vanuatu
French Polynesia: Society Is. - Tahiti, Passe de Papeete, reef sédiment, 3 spms, 01/1979 [Trn]; Tahiti, Arue, Chanel, lagoon, 12 spms, 12 m [Ltn|; Tahiti,
Mahina, Pointe Vénus, brushing on eoral rocks, I spm, 11/1989 [Trn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn]; Tuamotu Is. - Makemo, Pohuc, I spm, 83 m [Ltn]; Anaa, sédiment on external beach, 2 spms, 07/1980 [Trn]; Austral Is. - Rurutu, sandy sédiment near reef, 51m [Ltn]; Raivavae, sandy sédiment, lagoon, 2 spms [Ltn].
Horologica jayi Cecalupo & Perugia, 2012
Type locality: Mactan Is., Philippines.
French Polynesia: Society Is. - Tahiti, Papara, Mahaiatea, reef blocks, 1 spm, 14/07/81 [Trn],
Horologica micaelae Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Austral Is., Rapa - stn 10, Pointe Komire, + 79 m, 1 spm, 16-18 m, 27°34,8’S, 144°22,8”W, heap of stones covered with brown algae, 07 nov 2002; stn 28, Pointe Taekateke, 1 spm, 30 m, 27°38,4’S, 144°20,6’W, lot of stones covered with algae, 15 nov 2002; stn 43, Baie Ahurei, 3 spms, 45 m, 27°36,8’S, 14,4°18,3’W, drop off with muddy bottom at the base, 26 nov 2002; stn 44, NW Tauna Is., 12 spms, 30 m, 27°36,3’S, I44°18,2’W, drop off with muddy zones, 27 nov 2002; stn 47, Baie Ahurei, 5 spms, 33 m, 27°36,7’S, I44°19,rw, mud with eoral heads, 29 nov 2002; stn 48, off Pointe Rukuaga, 3 spms, 36 m, 27°34,rS, 144°22,rW, plateau covered by sand, 30 nov 2002.
Horologica nodosa Cecalupo & Perugia, 2012
Type locality: Bohol Is., Philippines.
French Polynesia: Society Is. - Tahiti, Papeete, Motu Uta, 2 m [Ltn]; Tahiti, Punaauia, outer slope, 20 m [Ltn]; Tahiti, Arue, outer slope, 15-60 m [Ltn]; Tahiti, Arue, lagoon, 10-18 m [Ltn]; Tahiti, Mahina, Baie de Matavai, 18 m [Ltn]; Tahiti, Mahaena, fringing reef [Ltn]; Tahiti, Tiarei, fringing reef, 1 m [Ltn]; Moorea, [Ltn]; Tetiaroa [Ltn], Mehetia, [Ltn].
Horologica paupercula Cecalupo & Perugia, 2012
Type locality: Bohol Is., Philippines.
French Polynesia: Society Is. - Tahiti, Taravao, Port Phaeton, in sand along beachfront, 3 spms [But].
Horologica tabauellii Cecalupo & Perugia, 2012
Type locality: Mactan Is., Philippines.
French Polynesia: Society Is. - Tahiti, Tiarei, sédiment, 1 spm, I m [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia, [Ltn].
18
A. Chcalupo & I. Pf.rugia
NOVAPFX 15(1): 1-22, 10 mars 20 14
Horologica gwenaellae Cecalupo & Perugia, 2013
Type locality: Espirilu Sanlo Is., Vaiuialu.
French Polynesia: Society Is. - Tahiti, Arue, outer slope, 20-40 m [Ltn]; Tahiti, Mahina, Les roses de Matavai, 45 m, 4 spms [LtnJ; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn],
Jociilator alliî'atiis Ceealupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Society Is. - Tahiti, Punaauia, outer slope, 20 m [Ltn]; Tahiti, Arue, outer slope, 15- 20 m [Ltn]; Tahiti, Arue, lagoon, 10-18 m [Lin]; Tahiti, Tiarei, fringing reef, 1 m [Ltn]; Tahiti, Ataahiti, sédiment, 3 spms, 30 m, 06/1985 [Trn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn],
Joculator ardiiinii Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Tuamotu Is. - Makemo, Pouheva, reef, 1 m. Passe Arikitamiro, 45 m [Ltn]; Ahe, sand on reef bord, 9 spms [Ltn],
Joculator carpatinus Cecalupo & Perugia, 2012
Type locality: Bohol Is., Philippines.
French Polynesia: Society Is. - Tahiti, Afaahiti, sédiment, 7 spms, 30 m, 06/1985 [Trn].
Joculator furvus Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Society Is. - Tahiti, Tiarei,
fringing reef, 1 m [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn].
Joculator inflatus Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Society Is. - Tahiti, Tiarei,
sédiment, I spm, 1 m [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn].
Joculator ianthinus Cecalupo & Perugia, 2013
Type locality: Espiritu Santo Is., Vanuatu.
French Polynesia: Austral Is. - Rapa, Hiri bay, 1 spm, 5 m [Ltn].
Joculator lividus Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Society Is. - Tahiti, Punaauia, outer slope, 20 m [Ltn]; Tahiti, Arue, outer slope, 15- 20 m [Ltn]; Tahiti, Arue, lagoon, 10-18 m [Ltn]; Tahiti, Tiarei, fringing reef, I m [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn].
Joculator minutus Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Society Is. - Tahiti, Afaahiti, sédiment, 13 spms, 60 m, 06/1985 [Trn],
Joculator recisus Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Society Is. - Tahiti, Mahina, Pointe Vénus, Banc du Dauphin, 2 spms, 18 m [Ltn]. Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn].
Joculator simulans Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Society Is. - Tahiti, Tiarei, sédiment, 1 spm, 1 m [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn].
Joculator suhconicus Cecalupo & Perugia, 2012
Type locality: Pamilacan Is., Philippines.
French Polynesia: Society Is. - Tahiti, Mitirapa, sédiment on the beach, 2 spms [But].
Joculator prunus Cecalupo & Perugia, 2013
Type locality: Espiritu Santo Is., Vanuatu.
French Polynesia: Society Is. - Tahiti, Arue, outer slope, 1 spm, 20 m [Ltn]; Tahiti, Tiarei, sédiment, 1 spm, I m [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn].
Joculator variahilis Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Society Is. - Bora Bora, in sédiment of the extreme point of the reef barrier, 2 spms [But].
19
A. Ci:iCALUPC) & 1. PliRUGlA
Cerithiopsidae and Newtoniellidae l'rom French Polynesia
Jociilator voucoseli Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia; Society Is. - Tahiti, outer slope, 1 spm, 65 m [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn].
Marshallopsis hlcmda Cecalupo & Perugia, 2012 Type locality: Pamilacan Is., Philippines.
French Polynesia: Society Is. - Bora Bora, in sédiment of the extreme point of the reef barrier, 2 spms [But];
Marquesas Is. - Nuku Hiva, sédiment, 1 spm, 10 m, 1980 [Trn].
Marshallopsis boucheti Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Society Is. - Tahiti, Passe de Papeete, reef sédiment, 1 spm, 06/1980 [Trn].
Marshallopsis gattellii Cecalupo & Perugia, 2012 Type locality: Bohol Is., Philippines.
French Polynesia: Society Is. - Tahiti, Papeete, Motu Uta, 2 m [Ltn]; Tahiti, Arue, outer slope, 15-60 m [Ltn]; Tahiti, Arue, outer slope, 22 spms, 20-57 m [Ltn]; Tahiti, Mahina, Pointe Vénus, Banc du Dauphin, 1 spm, 18 m [Ltn]; Tahiti, Mahina, Baie de Matavai, 18/20 m [Ltn]; Tahiti, Tiarei, fringing reef, 1 m [Ltn]; Tahiti Mahaena, [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn]; Tuamotu Is. - Rangiroa, near Passe de Tiputa, 7 spm, 100 m [Ltn].
Marshallopsis granosa Cecalupo & Perugia, 2012
Type locality: Bohol Is., Philippines.
French Polynesia: Society Is. - Tahiti, Tiarei, sédiment, 1 spm, 1 m [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn].
Marshallopsis limpida Cecalupo & Perugia, 2012
Type locality: Balicasag Is., Philippines.
French Polynesia: Society Is. - Bora Bora, in sédiment of the extreme point of the reef barrier, 2 spms [But].
Marshallopsis latea Cecalupo & Perugia, 2012 Type locality: Pamilacan Is., Philippines.
French Polynesia: Society Is. - Tahiti, Punaauia, blocks on the reef, 1 spm, 02/1982 [Trn]; Tahiti, Papara, ringing reef, I spm, 50 m, 06/1982 [Trn]; Tahiti, Papara, reef brushing, 2 spms, 0.50 m 30/11/1989 [Trn]; Tahiti, Arue, 1 spm, 30 m [Ltn]; Tahiti, Mahina, Orofara, sédiment on the beach, 3 spms [But]; Tahiti, Tiarei, sédiment, 1 spm, 1 m, [Ltn]; Bora Bora, in sédiment of the extreme point of the reef barrier, 1 spm [But]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn]; Marquesas Is. - Ua Pou, Haakuti, 2 spms, 70 m [Ltn].
Marshallopsis maesta Cecalupo & Perugia, 2012 Type locality: Balicasag Is., Philippines.
French Polynesia: Marquesas Is. - Ua Pou, Haakuti,
1 spm, 68 m [Ltn].
Marshallopsis melanesiana Cecalupo & Perugia,
2013
Type locality: Espiritu Santo Is., Vanuatu.
French Polynesia: Society Is. - Tahiti, Passe de Papeete, reef sédiment, 39 spms, 01/1981 [Trn]; Tahiti, Arue, 3 spms, 27 m [Ltn]; Tahiti, Arue, reef. Mission Poli Transect, 6 spms, 17/04/1982 [Tm]; Tahiti, Arue, sédiment on Lafayette reef, 1 spm, 1985 [Trn]; Tahiti, Mahina, Pointe Vénus, Banc du Dauphin, 18m, [Ltn]; Tahiti, Mahina, blocks on the reef, 1 spm, 02/1982 [Trn]; Tahiti, Mahina, Pointe Vénus, brushing on coral blocks, 1 spm, 11/1989, [Trn]; Tahiti, Mahina, Orofora, beach sédiment, 1 spm, 01/2009 [Trn]; Tahiti, Mahaiatea, blocks on the reef, 35 spms, 14/07/81 [Trn]; Tahiti, Afaahiti, sédiment, 52 spms, 30/60m 06/1985 [Trn]; Tahiti, Faaone, Tahiti, brushing on Coastal reef, 13 spms, 23/05/85 [Trn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn]; Tuamotu Is. - Anaa, sédiment on the externe beach, 5 spms, 1980-81 [Trn]; stn TH02-03, Hereheretue, 4 spms, 60 m, 19°52,48’S, 145°00,40’w, 28/04/2011, Meru 34; Moruroa Atoll, 6 spms [But]; Rangiroa, Passe de Tiputa, 25 spm, 81/100 m, [Ltn]; Makemo, Passe d'Arikitamiro, 6 spms, 45 m [Ltn]; Makemo, sédiment on reef front, 1 m [Ltn]; Anaa, sédiment on extern beach, 36 spms, 1980-81 [Trn]; Moruroa, sédiment on esterne beach, 14 spms, 1980 [Trn]; Austral Is. - Rurutu, sandy sédiment on ringing reef, 51 m [Ltn]; Raivavae, sandy sédiment on the lagoon, 2 spms [Ltn]; Banc du Président Thiers, N/O “Alis” campagne BENTHAUS, stn DW 1927, 3 spms, 105-95 m, 24°39’S, 146°01,6’W, 13/1 1/2002.
Marshallopsis tutuhaensis Cecalupo & Perugia, 2013
Type locality: Espiritu Santo Is., Vanuatu.
French Polynesia: Tuamotu Is. - Tikehau, in sédiment of the extreme point of the reef barrier, 1 spm [But].
20
A. Cecalupo & I. Perugia
NOVAI’I'X 15(1): 1-22, K) mars 20 14
MendaxcW mar^inata Siiler, 1908
Type locality: Three Kings Is., New Zealand.
French Polyncsia: Society Is. - Tahiti, Arue, outer slope, 6 spms, 35 m [Ltn]; Tahiti, Mahina, Baie de Matavai, 16 m [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn],
Mendax nifulus Cecalupo & Perugia, 2013
Type locality: Espiritu Santo Is., Vanuatu.
French Polynesia: Society Is. - Bora Bora, on sédiment on the outer slope of reef, 2 spm [But]; Tuamotu Is. - Moruroa, sédiment on the externe beaeh, 2 spms, 1980 [Trn],
Ondidopsis anime Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Society Is. -Tahiti, 1 spm, 20 m [Ltn]; Tahiti, Punaauia, outer slope, I spm, 30 m [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn]; Austral Is. - Rapa, Baie de Ahurei [Ltn]; Rurutu, sandy sédiment on fringing reef, 1 spm [Ltn];
Ondidopsis intricata Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Marquesas Is. - Ua Pou, Haakuti, 68m, 4 spms [Ltn].
Seda vaniiatensis Cecalupo & Perugia, 2013
Type locality: Espiritu Santo Is., Vanuatu.
French Polynesia: Tuamotu Is. - Anaa, sédiment on the externe beaeh, 1 spm, 07/1980 [Trn].
Syndiopsis alhachiarae Cecalupo & Perugia, 2012
Type locality: Bohol Is., Philippines.
French Polynesia: Society Is. - Tahiti, Papeete, Motu Uta, 2 m [Ltn]; Tahiti, Arue, outer slope, 15-60 m [Ltn]; Tahiti, Mahina, Baie de Matavai, 18 m [Ltn]; Tahiti, Mahaena [Ltn]; Tahiti, Tiarei, fringing reef, I m [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn].
Syndiopsis hongiardinoi Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Society Is. - Tahiti, Taravao, Baie de Port Phaeton, sédiment on the beaeh, 2 spms [But]; Tahiti, Mahina, Pointe Vénus point, brushing on coral blocks, 1 spm, 1 1/1989 [Trn].
Synthopsis cehiiensis Cecalupo & Perugia, 2012
Type locality: Cebu Is., Moalboal, Philippines.
French Polynesia: Society Is. - Tahiti [Ltn]; Motu One, sandy sédiment, 2 spms, I m, bord récif [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn].
Syndiopsis noninii Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Society Is. - Tahiti, Tiarei,
sédiment, 1 spm, 1 m [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn].
Synthopsis panglaoensis Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Society Is. - Tahiti, Tiarei,
sédiment, 1 spm, 1 m [Ltn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn].
Syndiopsis prima Cecalupo & Perugia, 2012 Type locality: Panglao Is., Philippines.
French Polynesia: Society Is. - Tahiti, Mahina, coral blocks on the reef, 2 spms, 16/05/82 [Trn].
Synthopsis praeaciita Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Society Is. - Tahiti, Papeete, reef sédiment, 2 spms, 06/1980 [Trn]; Tahiti, Punaauia, outer slope, 20 m [Ltn]; Tahiti, Arue, outer slope, 15- 20 m [Ltn]; Tahiti, Baie de Arue 10-18 m [Ltn]; Tahiti, Mahina, Pointe Vénus, brushing on coral blocks, 1 spm, 1 1/1989 [Trn]; Tahiti, Tiarei, sédiment,
1 spm, 1 m [Ltn]; Tahiti, Mahaena, fringing reef, 1 m [Ltn]; Tahiti, Mahaiatea, blocks on the reef, 2 spms, 14/07/81 [Trn]; Tahiti, Eaaone, brushing on the Coastal reef, 4 spms, 23/05/85 [Trn]; Tahiti, Afaahiti, sédiment, 3 spms, 60 m, 06/1985 [Trn]; Tahiti, Afaahiti, sédiment, 7 spms, 30 m, 06/1985 [Trn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn]; Marquesas Is. - Ua Pou, Hakahetau, 5m [Ltn]; Ua Pou, Haakuti 68 m, 4 spms [Ltn].
Synthopsis sdviae Cecalupo & Perugia, 2012
Type locality: Panglao Is., Philippines.
French Polynesia: Society Is. - Tahiti, Afaahiti, sédiment, 1 spm, 60m, 06/1985 [Trn].
21
A. Cecalupo & 1. Perugia
Cerithiopsidae and Newtoniellidae from French Polynesia
Spécula albengai Cecalupo & Perugia, 2013
Type locality: Espiritu Santo Is., Vanuatu.
French Polynesia: Austral Is. - Raps, Baie Hiri, 1 spm, 5 m [Ltn].
Spécula dubia Cecalupo & Perugia, 2013
Type locality: Espiritu Santo Is., Vanuatu.
French Polynesia: Society Is. - Tahiti [Ltn]; Bora Bora, reef, 2 spms, 1 m [Ltn]; Moorea [Ltn]; Tetiaroa, sandy sédiment [Ltn]; Mehetia [Ltn].
Spécula moalboalensis Cecalupo & Perugia, 2012
Type locality: Cebu Is., Moalboal, Philippines.
French Polynesia: Society Is. - Tahiti, Papeete, Motu Uta, 2 m [Ltn]; Tahiti, Arue, outer slope, 15-60 m [Ltn]; Tahiti, Mahina, Baie de Matavai, 18 m [Ltn]; Tahiti, Mahaena, [Ltn]; Tahiti, Tiarei, fringing reef, Im [Ltn]; Tahiti, Mahina, Pointe Vénus brushing on coral blocks, 4 spms, 0.50 m, 1 5/1 1/89 [Trn]; Tahiti, Afaahiti, sédiment, 5 spms, 30m, 06/1985 [Tm]; Tahiti, Afaahiti, sédiment, 3 spms, 60 m, 06/1985 [Trn]; Moorea [Ltn]; Tetiaroa [Ltn]; Mehetia [Ltn].
Tubercliopsis miranda Cecalupo & Perugia, 2012
Type locality: Pamilacan Is., Philippines.
French Polynesia: Society Is. - Tahiti, Afaahiti, sédiment, 1 spm, 60m, 06/1985 [Tm].
ACKNOWLEDGEMENTS
We would like to thank the malacological staff of the Muséum national d’Histoire naturelle of Paris, Philippe Bouchet, Virginie Héros and Philippe Maestrati, for the availability of their material and the scientific and bibliographie support; Michel Boutet (Tahiti), Jean Letourneux (Tahiti) and Jean Trôndlé (France), who hâve put their collections avaible to us.
We are also very gratefui to the Museo di Storia Naturale di Milano, particularly to Michèle Zilioli, for the SEM photos and to Elio Robba (UBM of Milano) for his constant suggestions. Thanks also to Jean Trôndlé, La Force, France, for his useful comments on the manuscript.
REFERENCES
Bouchet, P. & Rocroi, J-P. 2005. Classification and Nomenclator of Gastropod Families. International Journal of Malacology, Malacologia, 47(1-2): 1- 397.
Cecalupo, A. & Perugia F, 2012. Family
Cerithiopsidae H. Adams & A. Adams, 1853 in the Central Philippines (Caenogastropoda: Triphoroidea). Quaderni délia Civica Stazione Idrobiologica di Milano, 30[201 1]; 1-262. Cecalupo, A. & Perugia L, 2013. The Cerithiopsidae (Caenogastropoda, Triphoroidea) of the Espiritu Santo - Vanuatu- South Pacific Océan. Published by the authors, 253 pp.
Hutton F.W., 1885. New Species of Tertiary Shells. Art. LVl, Transactions and Proceedings of the Royal Society of New Zealand, 18: 333-335.
Jay, M. & Drivas, J. 2002. The Cerithiopsidae (Gastropoda) of Reunion Island (Indian Océan). Novapex, 3{\): 1-45.
Laseron, C.F., 1956. The Family Cerithiopsidae (Mollusca) from the Solanderian and Dampierian Zoogeographical Province. Australian Journal of Marine and Freshwater Research, 1 ( 1 ): 151-1 82, Figs. 1-57.
Marshall, B. A. 1978. Cerithiopsidae (Mollusca: Gastropoda) of New Zeland, and a provisional classification of the family. New Zealand Journal ofZoology, 5: 47-120, Figs 1-19.
Severns, M., 2011. Shells of the Hawaiian Islands.
The Sea Shells. ConchBooks, Hackenheim, 564
pp.
Trôndlé, J. & Boutet, M. 2009. Inventory of Marine Molluscs of French Polynesia. Atoll Research Bulletin, No. 50, National Muséum Natural History Smithsonian Institution, Washington. 87
pp.
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NOTE AUX AUTEURS
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Powell, A.W.B. 1979. New Zealand Mollusca. Marine, landand freshwater shells. William Collins Publishers Ltd: xiv + 500 pp.
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Keen, A.M. & Campbell, G. B. 1964. Ten new species of Typhinae (Gastropoda : Muricidae). TheVeliger7(\): 46-57.
Powell, A.W.B. 1979. New Zealand Mollusca. Marine, land and freshwater shells. William Collins Publishers Ltd: xiv + 500 pp.
Mayr, E. 1989. Attaching names to objects. In: What the philosophy of biology is : essaye for David Hull {M. Ruse, ed.), Klumer Academie, Dordrecht: 235-243.
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or by e-mail: vilvens.claude@skynet.be
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Novapex / Société 15(1), 10 mars 2014
1
N9VAPEX
Lieu de réunion : A partir de 14h, à notre local habituel :
Salle "Memling” (1er étage - ascenseur) - Rue de Genève, 470b - Schaerbeek (Bruxelles)
/ SoGiétt
Prochaines activités de la SBM
Claude VILVENS
SAMEDI 10 MAI 2014
Claude Vilvens : La détermination des escargots terrestres de Wallonie-Bruxelles
Que les naturalistes peu au fait de la malacologie éprouvent des difficultés à déterminer les escargots qu'ils rencontrent, spécialisés qu'ils sont en oiseaux, fleurs, champignons ou insectes, on peut le comprendre. Mais il n'est pas rare non plus de rencontrer des malacologues confirmés peu à l'aise dans ce genre d'exercice, simplement parce que leurs centres d'intérêt et de compétences se situent dans le monde des mollusques marins. Et on peut bien les comprendre puisque le président actuel de la SBM s'est d'abord spécialisé dans les Troques ;-) (donc des Marins), pour ensuite se dire qu'il n'était pas très normal de tout ignorer (ou presque) des mollusques terrestres de Belgique et d'Europe. Cela fait maintenant 20 and qu'il parcourt la campagne à leur recherche, seul ou avec les quelques autres membres de la SBM qui sont eux aussi des passionnés des "escargots". C'est une synthèse de ces observations et de cette expérience acquise qui est proposée ici à tous ceux qui ont envie d'en savoir plus sur les Clausilies, Moines, Maillots, Bulimes, Ambrettes, Hélices : bref sur nos "Escargots" !
SAMEDI 24 MAI 2014
Tout le monde : L'excursion de printemps de la SBM.
Avec le beau temps revient l'envie d'aller sur le terrain ;-) (air connu !). Nous envisageons de prospecter dans la région de Jette, dans quelques zones à haute valeur de biodiversité locale. Cette balade malaco sera aussi l'occasion de tester sur le terrain les coimaissances acquises lors de la conférence du 10 mai.
Durée de l'excursion : la matinée (rendez-vous à 9h30 - fin vers 12h30).
Comme d'habitude, les dernières informations seront disponibles sur notre site Internet (http://www.societe-belge-de- malacologie.be/) ou auprès de Claude (vilvens.claude@skynet.be) et d'Etienne Meuleman (e.meuleman@skynet.be). Comme d'habitude aussi, il convient de prévoir d'emporter sa bonne humeur, un guide de détermination ... et sans doute aussi bottes et vêtements de pluie (en principe, il fera doux et ensoleillé, mais bon ;-) . . .).
SAMEDI 14 JUIN 2014
Tout le monde : Le quiz : le grand jeu des coquillages
Avant les grandes vacances, nous vous proposons à nouveau de nous retrouver pour une activité ludique puisque nous allons en effet à nouveau jouer - vous l'aurez compris, il s'agira du quiz malacologique, formule qui a remporté beaucoup de succès lors de ses précédentes éditions, au point d'ne devenir une tradition pour fêter le début de l'été. Et on gagnera encore une fois quelque chose : mais oui, des coquilles bien sûr © Un joyeux après-midi avant les grandes vacances !
***
Réservez déjà dans vos agendas le 6/9/2014 et le 27/9/2014 (excursion).
Pour les informations de dernière minute :
f'.
Sur Internet :
http://www.societe-belge-de-inalacologie.be/
2
Nov APEX / Société 15(1), 10 mars 2014
Novapex/Société : la publication généraliste de la SBM
Rédacteurs en ehef ; Claude Vilvens & Etienne Meuleman
Tous les articles généraux sont les bienvenus pour Novapex/Société © !
Afin de faciliter le travail de la Rédaetion, il est vivement (et le mot est faible ;-)) souhaité de respeeter les règles suivantes pour les articles proposés :
♦ document MS-Word (pour PC Windows);
♦ police de caraetères Times New Roman;
♦ texte de taille 10, titres de taille 12;
♦ interligne simple;
♦ toutes les marges à 2,5 cm;
♦ document en une seule section;
♦ pas de mode colonne;
♦ photos en version éleetronique JPG ou PNG.
Merci pour les Scribes ;-) ! N'hésitez pas à demander une page avee en-tête pour cadrer au mieux vos travaux (vilvens. elaude(^skynet.be ou e.meuleman(@skynet.be).
Novapex/ Société 15(1), 10 mars 2014
3
Inventaire des mollusques présents dans le tube digestif d’un Spatangus purpureus O.F. Müller, 1776 à Favignana
(Iles Egades - Sicile - Italie)
Christiane DELONGUEVILLE
Avenue Den Doom, 5 - B - 1180 Bruxelles - christiane.delongueville@skvnet.be
Roland SCAILLET
Avenue Franz Guillaume, 63 - B - 1140 Bruxelles - scaillet.roland@skvnet.be
MOTS-CLEFS Echinoidea - Spatangus purpureus - Contenu digestif - Mollusques KEY-WORDS Echinoidea - Spatangus purpureus - Digestive content - Molluscs
RÉSUMÉ
Le contenu du tube digestif d’un oursin irrégulier, Spatangus purpureus, pêché au large de la Sicile, dans les Iles Egades, a été prélevé pour analyse. Celle-ci s’est limitée à l’inventaire des mollusques présents dans le sédiment ingéré par l’échinoderme et a révélé la présence de 23 espèces de gastéropodes, 21 espèces de bivalves et 1 espèce de polyplacophores pour un total de plus de 80 individus.
ABSTRACT
The gut content of an irregular sea urchin Spatangus purpureus, collected off the Egadi Islands in Sicily was taken for analysis. This was limited to an inventory of the molluscs présent in the sédiment ingested by the echinoderm. The analysis showed the presence of 23 species of gastropods, 21 species of bivalves and 1 species of polyplacophoran for a total of more than 80 individuals.
INTRODUCTION
Spatangus purpureus O.F. Müller, 1776 (Echinodermata - Spatangidae - Fig. 1) est un oursin irrégulier d’une douzaine de centimètres à symétrie bilatérale secondaire dont la bouche se situe sur la partie avant de sa face ventrale qui est plate et dont l’anus débouche sur l’arrière de sa face dorsale qui est bombée. Il est couvert de nombreux piquants, petits, fins et soyeux et de plus rares piquants allongés et plus solides implantés de façon régulière et symétrique sur sa face dorsale. Il est de couleur pourpre, ce qui est à l’origine de son nom. On le trouve dans l’Atlantique Nord-Est, de la Norvège au nord de l’Afrique ainsi qu’en Méditerranée (Hayward & Ryland 1998). Il vit sur des fonds sablo-graveleux ou détritiques depuis une faible profondeur jusqu’à généralement 70 à 80 mètres et même jusqu’à 900 mètres (Gage 1965). Il s’enfonce dans le sable et évolue à moitié enfoui lorsqu’il se nourrit en labourant le fond. Il participe ainsi au mélange et à la migration des sédiments du fond marin vers des zones plus profondes du sol, permettant ainsi à de petits métazoaires benthiques (méiofaune) de se nourrir. On le qualifie de détritivore car il se nourrit de matières organiques extraites des sédiments qu’il ingère. Barberà et al. (201 1) ont récemment étudié l’origine de la nourriture de Spantangus purpureus dans la région des Iles Baléares (Espagne). Sur base du type d’acides gras isolés dans les gonades, ces auteurs ont démontré que ces oursins sont omnivores et se nourrissent de débris d’algues lorsque leur milieu en est pourvu ou de résidus animaux et bactériens lorsque leur milieu est pauvre ou dépourvu en algues. Biagi (1975) s’est intéressé au contenu intestinal de Spatangus purpureus et Brissus unicolor (Leske, 1778) (un autre oursin irrégulier fréquent en Méditerranée) dragués par 50 à 70 mètres de profondeur dans l’Archipel Toscan. En pratiquant une ouverture dans la face dorsale de l’oursin l’auteur a observé le tube digestif contenant les sédiments ingérés par l’animal. Cet organe occupe la quasi totalité de la cavité. Un spécimen de Spatangus purpureus de taille moyenne (9x8 cm) peut contenir jusqu’à 90 g de sédiments en poids sec. Ces sédiments sont composés d’un mélange de sable, de débris d’algues coralligènes, de foraminifères, de fragments de test de crustacés ou d’autres oursins et de mollusques. Pour ces derniers, il s’agit généralement de fragments de coquilles (gastéropodes et scaphopodes), de valves dépareillées de bivalves et de plaques isolées de polyplacophores. Il est courant de rencontrer des gastéropodes juvéniles entiers et même des petits bivalves encore pourvus de parties molles, ceux-ci ayant été pris vivants lors de l’ingestion des sédiments par l’oursin. Une liste de 53 gastéropodes, 36 bivalves et 3 scaphopodes a été établie sur l’ensemble des spécimens disséqués.
Cet oursin est souvent accompagné de Montacuta substriata (Montagu, 1808) (Montacutidae - Fig. 4), petit bivalve inéquilatéral de forme ovale, dont l’umbo est placé dans la partie postérieure de la coquille. Sa taille ne dépasse généralement guère 4 mm. Il est caractérisé par la présence de stries radiaires sur chacune de ses valves. Il s’attache préférentiellement (Fig. 3) par un byssus à l’un ou l’autre piquant de la partie arrière ou ventrale du spatangue (Delongueville & Scaillet 2004).
4
Novapex / Société 15(1), 10 mars 2014
RÉCOLTES PERSONNELLES
Du matériel provenant d’un fond détritique (à 80 mètres de profondeur) a été collecté par une barque de pêche à Favignana (Iles Egades - Sicile) au mois de juin 2013 (Carte 1).
Dans le sédiment, les échinodermes étaient présents en très grand nombre : holothuries, ophiures [Astrospartus mediterraneus (Risso, 1826)], astéries {Astropecten aranciacus (Linnaeus, 1758), Hacelia attenuata Gray, 1840)] et oursins [Cidaris cidaris (Linnaeus, 1758), Spatangus purpureus O. F. Millier, 1776)]. Ceci explique la présence dans le matériel collecté de nombreuses espèces d’Eulimidae (gastéropodes parasites d’échinodermes), parmi celles-ci Sabinella piriformis Brugnone 1873, Nanobalcis nana (Monterosato, 1878), Parvioris ibizencus (Norsieck, 1968), Melanella frielei (Jordan, 1895), Sticteulima jejfreysiana (Brusina, 1869) et bien d’autres encore. Des spécimens vivants de Montacuta substriata (bivalves - Montacutidae) faisaient aussi partie de l’échantillon. Malheureusement, ni ce bivalve, ni les Eulimidae n’ont été trouvés fixés à leur hôte. Un spécimen de Spatangus purpureus non brisé de 10,0 x 9,0 cm a été isolé pour inventorier le contenu de son tube digestif. Une fois séché, l’oursin a été vidé et son contenu examiné à la loupe binoculaire. Il s’agissait essentiellement de fragments détritiques pouvant mesurer Jusqu’à 7 mm de long, parmi lesquels étaient présentes 23 espèces de gastéropodes, 21 espèces de bivalves et 1 espèce de polyplacophores pour un total de plus de 80 individus (Tableaux 1, 2 et 3).
Tableau 1
Anatomidae
Caecidae
Cerithiidae
Creseidae
Epitoniidae
Eulimidae
Lottiidae
Murchisonellidae
Raphitomidae
Rissoidae
Scissurellidae
Triphoridae
Trochidae
Turbinidae
Turritellidae
Vanikoridae
Gastropoda
Anatoma micalii Geiger, 2012 Caecum subannulatum de Folin, 1870 Caecum trachea (Montagu, 1803)
Bittium lacteum (Philippi, 1836)
Bittium submamillatum (de Rayneval & Ponzi, 1854) Creseis acicula Rang, 1828 Epitonium clathrus (Linnaeus, 1758)
Eulimidae sp.
Melanella frielei (Jordan, 1895)
Tectura virginea (O. F. Müller, 1776)
Ebala nitidissima (Montagu, 1803)
Raphitoma echinata (Brocchi, 1814)
Raphitoma sp.
Obtusella intersecta (S. Wood, 1857)
Pusillina inconspicua (Aider, 1 844)
Pusillina sp.
Scissurella costata d'Orbigny, 1824 Metaxia metaxa (Delle Chiaje, 1828)
Monophorus thiriotae Bouchet, 1 985 Jujubinus tumidulus (Aradas, 1846)
Bolma rugosa (Linnaeus, 1767)
Turritella commuais Risso, 1826 Megalomphalus azoneus (Brusina, 1865)
n
1
2
2
3 fragments 2 1 1 1 2 2 1
1 fragment
1 fragment 2 1 1 2 1
1 fragment 2 2
1 fragment 4,5 mm 1
Novapex / Société 15(1), 10 mars 2014
5
|
Tableau 2 |
Bivalvia |
n |
|
Arcidae |
Bathyarca pectunculoides (Scacchi, 1835) |
1 valve |
|
Astartidae |
Astarte sp. juvénile |
1 double |
|
Digitaria digitaria (Linnaeus, 1758) |
3 valves |
|
|
Goodallia triangularis (Montagu, 1803) |
1 double |
|
|
Carditidae |
Centrocardita aculeata (Poli, 1795) |
1 valve |
|
Coripia corbis (Philippi, 1836) |
2 doubles |
|
|
Kelliidae |
Kellia suborbicularis (Montagu, 1803) |
2 valves |
|
Limidae |
Limatula gwyni (Sykes, 1903) |
2 valves |
|
Limatula subovata (Monterosato, 1875) |
1 valve |
|
|
Leptonidae |
Hemilepton nitidum (Turton, 1822) |
1 valve |
|
Montacutidae |
Mancikellia parrussetensis (Giribet & Penas, 1999) |
2 valves |
|
Myidae |
Sphenia binghami Turton, 1822 |
1 valve |
|
Mytilidae |
Gregariella semigranata (Reeve, 1858) |
1 valve |
|
Modiolula phaseolina (Philippi, 1844) |
1 valve |
|
|
Neoleptonidae |
Neolepton sulcatulum (Jeffreys, 1859) |
8 valves |
|
Nuculidae |
Nucula nitidosa Winckworth, 1930 |
3 valves - 7 mm |
|
Propeamussiidae |
Parvamussium fenestratum (Forbes, 1844) |
1 valve |
|
Similipecten similis (Laskey, 1811) |
+ de 10 valves |
|
|
Psammobiidae |
Gari costulata (Turton, 1822) |
1 valve |
|
Semelidae |
Abra a/ba (W. Wood, 1802) |
1 valve |
|
Veneridae |
Timoclea ovata (Pennant, 1777) |
2 doubles |
|
Tableau 3 |
Polyplacophora |
n |
|
Callochitonidae |
Callochiton septemvalvis (Montagu, 1803) |
8 plaques |
DISCUSSION
Cet inventaire ne donne qu’une idée partielle de la composition de la faune malacologique présente sur ce fond détritique.
Les mollusques présents dans le contenu intestinal de l’oursin ne font pas l’objet de sa part d’une collecte sélective. En se déplaçant, il ingère tout simplement du sédiment dans lequel le hasard fera que se trouvent ou non des mollusques morts ou même vivants. La taille de ceux-ci est limitée par celle de l’ouverture de la bouche qui n’est pas extensible, puisque le test de l’oursin est rigide. L’ouverture buccale du Spatangus purpureus de Favignana est de 15,0 x 7,0 mm (Fig. 3). L’oursin ne peut donc avaler que des éléments dont la taille est inférieure à celle de l’ouverture buccale. Cela laisse néanmoins la place pour l’ingestion de particules détritiques de tailles relativement grandes, certaines valves de Nucula nitidosa présentes dans l’animal étudié mesuraient jusqu’à 7 mm. 11 n’est donc pas étonnant que les espèces de mollusques dont les adultes sont naturellement de petite taille puissent être ingérées sans problème {Caecum, Scissurella, Obtusella, Ebala, Pusillina, Hemilepton, Neolepton...) et même vivants {Caecum subannulatum avec son opercule, Goodallia triangularis et Coripia corbis encore avec parties molles). Pour les espèces dont les adultes atteignent une plus grande taille, leur présence sera limitée à l’ingestion de spécimens juvéniles uniquement {Bolma, Epitonium, Jujubinus, Turritella, Gari, Abra, Timoclea...), certains aussi pouvant être vivants {Timoclea ovata avec parties molles).
Ceci différencie le mode alimentaire passif de Spatangus purpureus du mode alimentaire actif de l’étoile de mer Astropecten aranciacus. Pour sa part, elle agit comme un prédateur qui effectue des collectes sélectives. Les mollusques font partie intégrante de son régime alimentaire. Grâce à sa grande taille (envergure 40 cm, diamètre du disque 10 cm), à la possibilité d’évagination de son estomac et à la relative plasticité de son ouverture buccale elle est à même de capturer des mollusques relativement grands.
6
Novapex / Société 15(1), 10 mars 2014
CONCLUSIONS
A l’instar de certaines étoiles de mer dont l’estomac peut contenir des mollusques, des oursins irréguliers peuvent également en contenir dans leur tube digestif. Pour les premières, les mollusques sont des proies et ils aboutissent dans l’estomac de l’échinoderme de façon active, car les animaux exercent une prédation volontaire. Pour les seconds, ils ne font pas partie de leur régime alimentaire, c’est le hasard qui décidera de leur présence dans le sédiment ingéré par les animaux pour trouver leur nourriture. Dans les étoiles de mer prédatrices de mollusques on peut trouver des coquilles de très grandes tailles, dans les oursins irréguliers, la taille des mollusques est fonction de la grandeur de l’ouverture de la bouche. Comme elle est étroite et rigide, l’oursin ne contiendra que des espèces de petites tailles ou des spécimens juvéniles.
Ces deux types d’échinodermes offrent une manière aisée de se procurer sur des bateaux de pêche des espèces de mollusques ou micro-mollusques benthiques.
RÉFÉRENCES
Barbera, C., Fernândez-Jover, D., Lôpez Jiménez, J.A., Gonzalez Silvera, D., Hinz, H., Moranta, J. 2011. Trophic Ecology of the Sea Urchin Spatangus purpureus Elucidated by Gonad Fatty Acids Composition Analysis. Marine Environmental Research', 71(4):235-246.
Biagi,V. 1975. Tanatocenosi di molluschi nel contenuto intestinale degli echinoidi irregolari Brissus unicolor (Leske) e Spatangus purpureus (O. F. Millier). Conchiglie', 1 1(7-8):149-164.
Delongueville, C. & Scaillet, R., 2004. Montacuta substriata (Montagu, 1808) vivant sur Spatangus purpureus (O. F. Millier, 1776) en Sardaigne. Novapex/Société', 5(4): 155-1 57.
Gage, J. 1965. Observations on the Bivalves Montacuta substriata and M. ferruginosa, “Commensals” with Spatangoïds. Journal of the Marine Biological Associationof the United Kingdom; 45:409 - 425.
Hayward, P.J. & Ryland, J.S. 1998. Handbook of the Marine Fauna of North-West Europe. Oxford University Press,
800 pp.
LÉGENDES
1. Spatangus purpureus - Favignana - Iles Egades (I) 10,0 x 9,0 cm. 2. Spatangus purpureus - Favignana - Iles Egades (I) - Ouverture buccale dégagée 15,0 x 7,0 mm. 3. Montacuta substriata sur Spatangus purpureus - Isola Rossa - Sardaigne (I). 4. Montacuta substriata - Isola Rossa - Sardaigne (I) 3,1 x 2,7 mm.
Novapex / Société 15(1), 10 mars 2014
7
L’écho des réunions
Christiane DELONGUEVILLE & Roland SCAILLET / Marc ALEXANDRE
Réunion du 14 décembre 2013 (MA et CV) ^ Roland Scaillet et Christiane Delongueville : Récoltes malacologiques en République turque de Chypre du Nord
C 'est presque devenu une tradition au sein de notre société de terminer l 'année en ayant le plaisir de donner la parole à nos deux globe-trotters passionnés de mollusques Européens. Ce rendez-vous attendu avec impatience par chacun d’entre nous est à chaque fois une superbe découverte que ce soit d’un point de vue géographie, historique, mais aussi et surtout malacologique. Mais trêve de balivernes laissons plutôt la parole à nos deux compagnons de voyage.
« La République turque de Chypre du Nord est située à l’est de la Méditerranée dans le bassin levantin. Des photos de monuments et de sites ineontoumables ont raconté l’histoire de l’île depuis le temps des Romains en passant par celui des Francs (les Lusignan), des Croisés, des Vénitiens et des Ottomans pour terminer avec l’Empire Britannique, l’indépendance de l’île et sa division actuelle en deux états autonomes.
Autour de l’île de Chypre, les courants marins de surface tournent dans le sens inverse des aiguilles d’une montre. La position de l’île dans le bassin levantin la place dans des eaux dont la salinité et la température sont plus élevées que celles relevées en moyenne dans les autres parties de la Méditerranée. Tous ces paramètres (courants, température et salinité) contribuent à en faire un endroit propice à l’installation d’espèces invasives, qu’elles soient d’origine lessepsienne ou dues au développement de la navigation en provenance de régions tropicales. Le nord de l’île est situé à quelques 70 km de la côte turque, à l’est, la Syrie est éloignée de plus de 100 km. L’éloignement relatif de l’île de Chypre des côtes libanaises et israéliennes peut contribuer à expliquer le nombre relativement faible d’espèces invasives répertoriées autour de l’île en comparaison de ces pays.
L’iconographie malacologique qui a été présentée était le fruit de trois voyages dans ce pays aux plages idylliques et désertes fréquentées par deux espèces de tortues marines qui viennent y pondre leurs œufs en toute tranquillité. Pour les mollusques, ont été illustrées des espèces pélagiques (Janthina janthina et Janthina prolongata), des espèces parasites d’oursins (Vitreolina philippi), des espèces classiques (Typhinellus labiatus collecté dans l’estomac d’une astérie, Astropecten aranciacus), des espèces de la Méditerranée orientale {Char onia variegata, Nassarius gibbosulus ou encore Nassarius circumcinctus) et enfin des espèces invasives (parmi d’autres : Septifer forskali, Acteocina mucronata, Cinguinla isseli...). Parmi les techniques de récolte présentées, les plus productives ont été la collecte de laisse de mer, le nettoyage des filets de pêche et la plongée en apnée dans les eaux peu profondes. Au total, 1 8 espèces invasives de mollusques marins (bivalves et gastéropodes confondus) sur les 42 répertoriées en 2009 dans la littérature ont été illustrées par des spécimens photographiés in situ ».
1. Septifer forskali 9,7 x 6,9 mm - 2. Brachidontes pharaonis 7,9 x 5,3 mm - 3. Rissoina bertholleti 5,2 x 2,3 mm 4. Cerithium scabridum 19,4 x 7,9 mm - 5. Ergalatax junionae 21,8 x 1 1,4 mm - 6. Aplysia dactylomela 200 mm 7. Dendrostrea frons 23.9 x 11,1 mm
Un grand merci à Roland et Christiane pour cet exposé riche en informations et en illustrations.
8
Novapex / Société 15(1), 10 mars 20 1 4
Du neuf du côté des publications de la SBM Les posters de la SBM déjà disponibles : format A3, plastifiés - 5.00 EUR + port.
Renseignements ; Etienne Meuleman - e.ineuleman(^skynet.be
Ç,i«o>j)i8ûac
AtbmâriM prscc/ara iL PfcilTffr 115)1 t } ifun • OrMve iCreu)
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; A/biaâTM puellM tt PletRei- IHîOi ' 15 I RSffûtec.oSitina»)
Principaux mollusques d’eau douce de Wallonie
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1 Va)l |
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Et deux nouveaux posters en ce début 2014
Novapex / Société 15(1), 10 mars 20 1 4
9
t Quelques nouvelles publications
Roland SCAILLET & Claude VIL VENS
Accrescimenti Tome V
Stadi di accrescimento dei molluschi marini del Mediterraneo
par Maria Scaperrotta - Stefano Bartolini - Cesare Bogi
1 92 pages
Editeur ; L’Informatore Piceno - Ancona - Italie - 2013
Prix : 65.00 EUR
Maria Scaperrotta Stefano Bartolini Cesare Bogi
ACCRESCIMENTI
STADI DI ACCRESCIMEN ï O DEI MOl^LUSC'HI MARINI DEL MEDITERRANEO
Stages of growth of'llic marine moîluses of the Mcctiterranean Sea
Volume V
lorsqu’on ne dispose pas d’une vue d’ensemble des espèces.
Et bien oui, voilà que vient d’être publié le cinquième volume d’un Opus commencé en 2009 :
« Accrescimenti - Stadi di accrescimento dei molluschi marini del Mediterraneo ». Les auteurs nous y annoncent déjà qu’ils travaillent à l’élaboration du sixième tome. Si nous l’attendons avec impatience, ne boudons pas notre plaisir et voyons d’abord ce que nous apporte le cru 2013.
126 espèces viennent s’ajouter à celles déjà illustrées. A ce jour, 632 espèces ont fait l’objet de ce concept unique: nous présenter pour chacune d’elles leurs différents stades de croissance. Cette entreprise n’est pas sans nous réserver des surprises, tant parfois et même souvent la forme des spécimens juvéniles diffère du tout au tout de celles des exemplaires adultes. Dans ce cinquième volume, on retiendra particulièrement l’illustration d’espèces invasives sur base de matériel en provenance des côtes d’Israël comme Cerithidium diplax, Cerithidium perparvulum, Diodora ruppellii, Electroma vexillum, Monotygma watsoni, Murex forskoehli, Odostomia lorioli, Timoclea roemeriana. Parmi les gastéropodes, une attention particulière a été consacrée à des éléments du zooplancton comme nombre de représentants de la famille des Atlantidae, des Cavoliniidae, des Cliidae et des Limacinidae. Les bivalves n’ont pas été oubliés, avec un effort tout particulier portant sur les familles des Limidae, Semelidae et Thraciidae. En fin de volume des planches rassemblant les espèces d’une même famille ont l’intérêt de souligner des différences qui bien souvent restent subtiles et difficiles à appréhender
Pour chaque taxon spécifique, la liste systématique des espèces illustrées à ce jour publiée en début d’ouvrage renvoie le lecteur au volume correspondant. Si l’Italien n’est pas votre langue maternelle, il n’y a aucun problème. La fiche technique se rapportant à chaque espèce est traduite en Anglais en fin de volume.
Digérons le volume V en tout premier lieu et attendons avec impatience ce que nous réservent Maria Scaperrotta, Stefano Bartolini et Cesare Bogi dans le sixième volume déjà en préparation.
Avis aux amateurs et bonne lecture.
Roland Scaillet
10
Novapex / Société 15(1), 10 mars 20 1 4
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Plancton, aux origines du vivant |
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par Christian Sardet |
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216 pages, 550 illustrations |
Editions Ulmer : |
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ISBN : 9782841386345 |
http;//ww'w.editions-ulmer.fr./livre/plancton-aux-oricines- |
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Année d'édition : 2013 |
du-vivant-430-l.htm Prix: 39.90 EUR |
Voici bien un livre magnifique !
D'abord par son iconographie, présentant les divers organismes du plancton dans leur milieu : les photos sont à couper le souffle, tant elles sont précises, lumineuses sur un fond sombre, parfois surprenantes, toujours d'une telle qualité que l'on peut parler de photo d'art.
Ensuite, par la richesse de l'information contenue : c'est à un véritable parcours exhaustif de toutes les composantes du plancton que le lecteur est invité.
Bien sûr, il y a notamment des mollusques, comme les céphalopodes, les nudibranches et surtout les ptéropodes et les hétéropodes ;
Mais il y a aussi des crustacés en tous genres (ostracodes, copépodes, amphipodes ...), des cnidaires (où, à côté des méduses classiques, on découvre que les eoraux calcaires ne sont pas les seules communautés : les siphonophores en constituent une autre forme), des éponges, des être unicellulaires, très nombreux et aux formes étranges (foraminifères, coccolithophores, diatomées, radiolaires)
A remarquer encore la présence de QR codes insérés dans les différents chapitres du livre : ils permettent d’accéder avec un smartphone ou une tablette aux 20 vidéos des "Chroniques du Plancton" (www.planktonchronicles.org/ff).
Un cadeau splendide à offrir ou plutôt à s'offrir ;-) !
Claude Vilvens
Novapex / Société 15(1), 10 mars 2014
Nous avons reçu
Etienne MEULEMAN
LES NATURALISTES DE LA HAUTE-LESSE
(Belgique)
N°274, novembre — décembre 2013
|
Calendrier des prochaines activités |
2 |
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Compte rendu des activités |
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Inventaire biologique de la mare de Sohier deux ans après le curage |
4 |
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Balade à Revogne-Froidiieu centrée sur les hirondelles et les vieux chemins communaux réaménagés |
6 |
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Journée d'ornithologie en Zélande |
8 |
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Relevé des mares à la réserve du Grand Quart? à Baronville |
10 |
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Sortie mycologique |
13 |
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Prospection de la Réserve de Saint-Rémy à Rochefort |
14 |
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Le Tchad et les changements climatiques |
23 |
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Chronique de l'Environnement |
27 |
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Informations |
29 |
XENQPHORA
(France)
N°142, avril-juin 2013
4 Le coin du Débutant par G. Jaux
7 Dernières trouvailles aux Fidji par Th. Vulliet
8 Découverte de Conus harthelemyi à Madagascar par F. Goûtai 10 Une étonnante variabilité par P. Bail
13 Note sur la présence de Indothais blanfordi (Melvill, 1 893)
au Sénégal et en Guinée-Bissau par R. Flouart et J-L. Delemarre
20 Hommage à notre Ami Michel Gueguen par D. Gratecap
21 Conus lamberti : une rectification
22 Curveulima dautzenbergi (Pallary, 1900) en Bretagne par C. Delongueville et R. Scaillet
27 Une classification généalogique des Xenophoridae par N. Laurenceau
37 Coquillages des îles Marquises par S. Pineau et J-P. Duboc 44 Reçu au Club par P. Bail
46 Informations du MNHN
47 Echo...quillages
48 Micro-mollusques par S. Aiken
12
Novapex / Société 15(1), 10 mars 2014
SPIRULA
(Pays-Bas)
N° 393, juillet-août 2013
Diverse bronncn Excursies en Malacologische agenda Nederland 109
H.P.M.G. Menkhorst Overpeinzingen op een zonnige voorjaarsmorgen 111
C.M. Neckheim Enkele vondsten van mariene mollusken uit Suriname, waaronder
Epitonium tollini Barlsch, 1938, die nieuw is voor de
fauna van Suriname 112
B. van der Valk De lagune van de Zandmolor tussen Kijkduin en Ter Heijde,
Zuid-Holland: een nieuw waddenhabitat voor tweekieppige mollusken
op de voorheen rechte Zuid-Hollandse kust 115
B. Langeveld & H. Mulder Een misvormde Mactra stuUorurn plistoneerlandica Van Regteren
Aliéna, 1937 van de Zandmotor (Zuid-Holland): sifo-grazende
platvissen in het Eemien 117
Nieuw in Nederland (samengesteld door G.D, Majoor) 119
G. Beuker Vondsten van de Noorse rotsboorder Panomya norvegica
(Spengler, 1793) van het Nedcrlands Continentaal Plat 119
W. Faber Weekdiercn op postzegels - Molluscs on stamps 120
W, Faber Nieuw beschreven mariene molluskensoorten - (new taxa:
marine molluscs) 122
W. Faber Artikelen in tijdschriften - (journal papers: marine malacology) 123
W. Faber Nieuwe boeken - new books 127
W. Faber Schelpenbeurzen en bijeenkomsten 128
MOLLUSC WORLD
(Grande-Bretagne) N°32,juin 2013
Contents
3 Conservation Ofîicer’s Report 2012 Martin WiUing
6 ‘On the spot’ questionnaire: Graham Long
Marine Recorder’s report 2012 Bas Payne
1 Non-marine recording 2012 Adrian Norris
9 Some recent snail records from Southern England
Jonty Denton
10 Specialist predators of molluscs C/ive Craik
12 Book: A Slow Passion by Ruth Brooks Peter Topley
13 A shelling trip to the north of Britain
John Llewellyn-Jones
14 TRGAVTKG forthcoming publication
15 The flame shells of Kyle Akin Dcw Harries
18 The snails of St Sebald Adrian and Barbara Norris
19 Nerites of the Matutinao River, Cebu, Philippines
Malcolm Symonds
20 British Shell Collector's Club meetings
22 Field meeting: South Pembrokeshire,
16"’-20"’ September 2012 Celia Pain
26 J. Wilfrid Jackson’s conchologicai correspondents
Brian Goodwin
28 Letter: Ciant African snails Sarah Lucas
29 Bon. Treasurer’s Report on the financial
statements to 31” December 2012 Nick Light
30 About the Society/Instructions to authors 32, 31 DIARY OF MEETINGS/ new members
Novapex / Société 15(1), 10 mars 2014
13
IBERUS
(Espagne)
Vol. 31,N° 2, 2013
Indice
Iberus 31 (2) 2013
Ryall R, Horro J. & RoiAn E. A révision of the genus Genota H. and A. Adams, 1853 (Gas- tropoda; Conoidea; Borsonidae) from West Africa
Révision del género Genota H. y A. Adams, 1853 (Gastropoda; Conoidea; Borsonidae) de
Africa Occidental G 1 7
Guerra a., Caro MaB., Seai.ey M.J. and Lozano Soedevilla F, Two new records of octopods in Ganary Islands; Amphioctopus burryi (Voss, 1950) and Macrotritopus defilippi (Vérany, 1851) [Cephalopoda: Octopodidae]
Dos nuevos registros de pulpos en las islas Canarias: Amphioctopus burryi fVoss, 1950) and Macrotritopus defilippi 1851) [Cephalopoda: Octopodidae] 19-26
Hoeyoak D.T., Hoeyoak G.A., Torres Alba J.S., da Gosta Mendes R.M. and Quinonero SaLGADO s. Succinea {Calcisuccinea) sp., an American land-snail newly established in Por- tugal and Spain (Gastropoda: Succineidae)
Succinea (Calcisuccinea) sp., un caracol terrestre americano nuevo para Portugal y Espana
( Gastropoda: Succineidae) 27-39
Rubio F., FernANDEZ-GarcÉS R. and RolAn e. The genus Haplocochlias (Gastropoda, Skenei- dae)
El género Haplocochlias (Gastropoda, Skeneidae) ,41-126
Sibaja-Cordero J.A., GarcIa-Méndez K. and TroncoSO J. S. Additions to the mollusk checklist of Cocos Island National Park, Costa Rica (Eastern Fropical Pacific)
Adiciones al catalogo de los moluscos del Parque Nacional Isla del Coco, Costa Rica (Pacifico Tropical Este) 1 TJ- 1 63
PelorCE J., Horst D. et Hoarau a. Une nouvelle espèce de la famille Aglajidae (Gastropoda: Opisthobranchia) des côtes de Méditerranée française
Una nueva especie de la familia Aglajidae (Gastropoda: Opisthobranchia) de la costa francesa del Mediterràneo 165-170
Notas brèves
GuaELART J., Acevedo 1., CalvO M. y MachORDOM a. Protocole no létal para la obtencion de muestras de tejido (para estudios genéticos) en la lapa amenazada Patella ferruginea (Gas- tropoda, Patellidae)
Non-lethal protocol for tissue-sampling (for genetic studies) in the endangered limpet Patella IcuugmtdL (Gastropoda, Patellidae) 171-174
14
Nov APEX / Société 15(1), 10 mars 20 1 4
SPIXIANA
(Allemagne)
Vol. 36, N°l,2013
INHALT - CONTENTS
Seite
Ermilov, S. G., D. Sandmann, F. Marian & M. Maraun: Two new oribatid mite species of
the genus Gittella from Ecuador (Acari, Oribatida, Oppiidae) 1-8
Ermilov, S. G. & W. Niedba+a; Contribution to the knowledge of the oribatid mite fauna of
Bolivia, Zambia, Cambodia and Vietnam, with descriptions of two
new species (Acari, Oribatida) 9-19
Fôrsterra, G., V. Hâussermann, R. R. Meizer & A. Weis: A deep water pycnogonid close
to the beach: Colossendeis macerrima Hoek, 1881 spotted at 18 m
in the Chilean fjords (Chelicerata, Pycnogonida, Colossendeidae) . 20
Langegger-Weinbauer, C. & L. v. Salvini-Plawen: Anatomical redescription of two species
of Philinoglossidae (Gastropoda, Cephalaspidea) 21-41
Uribe, R. A., K. Nakamura, A. Indacochea, A. S. Pacheco, Y. Hooker & M. Schrôdl: A review
on the diversity and distribution of opisthobranch gastropods from Peru, with the addition of three new records (Gastropoda, Hetero- branchia) 43-60
Ferrari, R. R., C. Schlindwein & A. Nemésio: Description of the female Euglossa perpulchra
Moure & Schlindwein, 2002 and an identification key to the females of the Euglossa decorata Smith, 1874 species group (Hymenoptera,
Apidae, Euglossina) 61-65
Horstmann, K.: Révisions of Nearctic Tersilochinae IV. Genus Phradis Fôrster (Hy- menoptera, Ichneumonidae) 67-92
Pfôciennik, M., P. Gadawski, D. Tempelman & H. W. Riss: First records from Roland of
Glyptotendipes ospe// Contreras-Lichtenberg & Kiknadze, 1999 and G.signatus (Kieffer, 1909) (Diptera, Chironomidae) 93-96
Jàger-Zürn, I., M. Spies, C. T. Philbrick, C. R Bove & A. Mora-Olivo: Plant galls (cecidia)
in the neotropical water plant family Podostemaceae induced by
larvae of Chironomidae (Diptera) 97-112
Wirtz, P, A. Brito, J. M. Falcôn, R. Freitas, R. Fricke, V. Monteiro, F. Reiner & O.Tariche: The
Coastal fishes of the Cape Verde Islande - new records and an annotated check-list (Pisces)... 113-142
Rakotoarison, A., J. Kôhler, F. Glaw & M. Vences: The advertisement call of the relict frog
Tsingymantis antitra from Madagascar (Anura, Mantellidae) 143-148
Darilmaz, M. C., Ü. incekara & R. Vafaei: Contributions to the knowledge of Iranien
Aquatic Adephaga (Coleoptera) 149-152
Baehr, M.: New species of the genus Pentagonica Schmidt-Goebel, 1846 from
the Oriental Région. Supplément to “The genus Pentagonica Schmidt-Goebel in the Oriental, Papuan, and Australien Régions”
(Coleoptera, Carabidae, Pentagonicini) 153-159
Buchbesprechungen 42, 66, 160
Erratum 1 60
Novapex / Société 15(1), 10 mars 2014
15
THE NAUTILUS
(U. S. A. - Californie) Vol. 127, N°2, juin 2013
|
CONTENTS |
Voit mie 127, Nimd'jer2 Julie 21, 2013 ISSN 0028-1344 |
|
|
Yuri I. Kantor |
Aiitarctica, whcre turrids and w'iielks converge: A rcxision of |
|
|
M.G. Harasevrych |
lùiLsiitu>linui Powell, 1951 (Neogastropoda: Buccinoidca) and a |
|
|
description ofa ncw genns |
43 |
|
|
Alisa Kosyan |
Aide» u^fiisiis i}^>u>tus new geniis and ncw spccies, a ncw bnccinid |
|
|
Yuri I. Kantor |
(GiLstropoda: Neogirstropcxla) froni tlic Nord» Pacific Occtui |
|
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Michael J. Bolton |
A new species of Striostreu (Bivalvia: Flemingo.streidae) frorn the |
|
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Roger W. Portell |
upper Pliocène and lowcr Pleistocene strata of Florida, USA |
65 |
|
G. Thomas Watters |
Rediscovery of Choanopoitm? smithianuin Pfeiffer, 1866 |
|
|
Jozef Grego |
(Annulariidae) from Haiti and désignation of a neotype, with the |
|
|
Jozef Stefîek |
description of two new species oi Weinlamlipoina Bartsch, 1946 |
78 |
|
Orso Angulo-Campillo |
Mariania kinoi (Nudihranchia: Tritoniidae): A new species |
|
|
Hans Bertsch |
from the tropical eastern Pacific |
85 |
|
Research Note |
||
|
Timothy A. Pearce |
Omplialotrnpis ilapinjensis , a replacement name for O. costulata |
|
|
Megan E. Paustian |
Emberton and Pearce, 1999 (Castropoda: Littorinimorplia: |
|
|
Assimineidae) |
90 |
THE FESTIVUS
(U.S.A.)
Vol. XXXXV, N°9, septembre 2013
Club news 76
Observation on the biology of the Chestnut Cowry Neobemaya spadicea in Mission Bay
PAULTUSKES 77
Notes on the cowry Neobemaya spadicea: novel observation of an unspotted and uniquely colored mantle
JUDITH LEA GARFIELD 81
The San Diego Shell Club Bazaar
LARRY BUCK & DAVID B. WALLER 83
Vol. XXXXV, N°10, octobre 2013
Club news 86
Some notes on prédation in marine gastropod shells on Sanibel island, Lee County, Florida
SUS AN HEWITT 87
Collecting Pismo Clams Tivela stultorum (Mollusca; V eneridae)
PAUL TUSKES & DAVID B. WALLER 90
16
Novapex / Société 15(1), 10 mars 20 1 4
AMERICAN CQNCHOLQGIST
(U.S.A. - Sud-Est)
Vol. 41, N° 3, septembre 2013
Editor's comments 3
Hâve you heard of Kiritimati? by Marcus Coltro 4
Révision of the Worldwide Recent Piiinidae and sonie remarks on fossil European Pinnidae reviewed by Thomas Eichhorst 9
Color variations in Muricopsis principensis Rolân &
Fernandes, 1991 (Gastropoda. Muricidae) from
Principe Island by Roland Houart & Sandro Gori 10
A day with Mitsuo Chino, President of the Tokyo Malacological Society by Kristina Joyce — 12
Dealer Directory 14
The Shell collection of His Majesty The Emperor Showa of Japan by Kristina Joyce 16
COA Convention 2013 byTom Eichhorst 18
2013 Neptunea Aw'ards by Harry Lee 21
Size matters: océan acidification and warming alter metabolic scaling in Polyplacophora by Nicholas Carey 22
Shell collecting as seen by Gene Everson
by Tom Eichhorst 24
An interesting find by Paul Kanner 27
COA Grant Donation Program 28
2013 COA Research Grants 28
In Memoriam 29
Sybil B. Burger by Tom Eichhorst 29
“Available” vs. “Valid” 29
COA 2014 - 11-15 August 2014, Wilmington, NC 30
BULLETIN OF MALACOLOGY
(Taïwan)
N°36, janvier 2013
1 . A New Species of Oocorys (Gastropoda rCassidac) from the Pratas Islands, near Taiwaa.. Kim Yang Lai. . . 1
2. New Record of Paphk alapapilionis Rôding 1 798 from Peng Hu, Taiwan
Chih Yimg Chen, Shu Chen Ho, Wen-Lung Wu. . .7
Paphia alapapilionis Rôding 1798 ' Æ:5CI^...7
3. Bbgeographical distribution of the endemic land snail Takasagohadra rauHifiasciata Kuroda, 1941
(Pulmonata: Biadybaenidae) with the description of a new species of genus Takasagohadra from China:
A new discovery between mainland China and Taiwan Island Shu-Ping Wu, Chung-Chi Hwang...l 1
+ — Takasagohadra Uni
' *tlt«...ll
4. ' aitaiS ' niFU '
Novapex / Société 15(1), 10 mars 2014
17
l liKNAND & RIKA DE DONDER Melsbroekscstraal 21 1800 Vilvoordc Pcutic BELGIUM
'Pel : +32 (0)2 253 99 54 Fax : + 32 (ü)2 252 37 15 e-mail ; fernand.d.e.dondcrC«^panclora.bc
WORLDWIDE SPECIMEN SHELÏM
10 Minutes froni Brussel.s AirporL. Visiilors wclcome.
Alî l amilics from lhe very coniirum u> lhe ultra rare, .spccialiî^cd in Pccrtinidao, Philippine shells and European ühclls.
P/'ee îist ou regriPAf, gt*od. quaîily shcîls at r/ic bc.st prireç. Sciti s factum ^uaTantcis.) !
The quarterly bulletin of the Conchological Society of Southern Africa contains reviews and discussion of Southern Ahrican marine and non-marine shells, and information about shell collecting in the région. Membership of the Society is US$25 per year.
Please contact
The Conchological Society of S.A.
7 Jan Booysen Str.
Annlin 0182 Pretoria South Africa or
email niikec(^insinfo.niintek.ac.za
Si vous passez commande chez l'un de nos annonceurs, n'oubliez pas de préciser que vous avez trouvé son annonce dans Novapex/Société !!!
SOCieOAD ESPAftOLA Ù£ MALACOLOGIA
il
Mubco Naclonal de Cienctas Naturalea José Gutiérrez Abascal, 2 28006 MADRID
SEM (Sociedad Espahola de Malacologia) is a scientic society devoted to lhe study of molluscs.
Every year the memberships reccive lhe following publications:
2 issues of IBERUS
1 issue ofRESEN AS MALACOLOGICAS 2-3 issues of NOTICIARIO DE LA SEM
some years, 1 extra IBERUS from a Congress or as a supplément.
You can be membership of the SEM by 7.000 ptas by year, plus an unique inscription fee of 1.000 ptas.
Please, ask for the inscription print paper.
XEMOftlORA
Française
2010 Yearlv subscriDtion rate
France -Eurone - DOM TOM : 50
Other countries : 60 €
Visit our site :
gÆ BP 307 F-75770 Paris (ledex 16
18
Novapex / Société 15(1), 10 mars 20 1 4
Nederl^)îsF"^/^' ^,Mdlâcologjsthe S— V^r^îiiging
f'
Dutch
Malacoiogical
Society
Our society warmly welcomes new members (both from the Netherlands and abroad) to participate in our activities:
- the journals (Basteria and Correspondentieblad)
- the meetings {usually 3-4 per year)
- the Internet website
- the library
- the collecting excursions
Juin us and niecl ncw shelling friends. Fiinher info: Brani Brcure, Van Schagcnplaiitsoen 8, NI.-2741 EN Waddinxveen, The Netherlands. E-mail; abreure@xs4all.nl
GLORIA MARIS
A magazine dcdicated to the .study of shell.s.
Edited by the Belglan Society for Conchology, organizers of the Belgium Shellshow
Subscription: Belgium: € 30 - The Netherlands: € 33 Other countries: € 40
Members account manager: ,1. W uyts Koningsarendlaan 82 B 2100 Belgium
tel.: 32 3 324 99 14 c-mail: wuyts.jeanrn scarlet.be
Club Conchylia
lui'' German Shell Collector's Club e.v
Our journals:
^ Conchylia # Mitteilungen
Acta Conchyliorum
Yeariy subscription rate: 50.- €
Visit our site:
www.club-conchylia.de
Further information:
Klaus Kittel
Sonnenrain iO
D-97859 Wiesthal
e-mall: klaus_kittel@hotmail.com
Kcp[)cl Bay dddings A quarterly magazine dedicated to the study of shells.
Fdited by the Keppel Bay Shell Club Inc. Subscription:- $20.00 Aus.
Apply to:- Keppel Bay Shell Club Inc. P.O. Box 5166
Central Queensland Mail Centre, 4702 Queensland,Australia.
Novapex / Société 15(1), 10 mars 2014
19
TRITON
Journal of the Israël Malacological Society
ISSN 1565-1916
Published twice a year since 2000 Yearly subscription rate 20 €
Further information:
Eduard Heiman
e-mail; heimel(gnetvision.net.il
I
ÈWltTliiH.t'lïPr
Calendar inembership (Jan - Dec) = $25 (USA) Postal surcharges: + $5 for USA first class, Canada & Mexico + $5, other nations + $15
New membcrs ajrply to üuris I nderwood, Mcnibership Director 698 Sheridan Woods Drive VV. Melbourne, FL 32904-3302 USA
dunderwoodl(« cfl.rr.com
Quarterly Journal of the Conchologists of America, Inc.
PHILLIP W.CLOVER TEL/FAX# 707 9% 6%0 P. O. BOX 339 cloversheUs@.t uno.com
GLEN ELLEN CA. 95442 USA
DEALER DSr WORLD WIDE SPECIMEN SEASHELLS SINGE 1960,SPECIALIZING IN RARE & COMMON CYPRAEA, CONUS,
VOLUTA, MUREX, MITRA, EPITONIUM,LATIAXIS, OVULA PLEUROTOMARIA,PECTENS,ETC. ALSO CURRENT AND OUT OF PRINT SHELL BOOKS. free price lists on reouest
Calamar géant dans le Pacifique
TOKYO Des scientifiques et des chaînes de télévision japonaise et américaine ont annoncé hier avoir filmé pour la première fois un cala- mar géant par 900 m de fond dans l’océan Paci- fique. L’animal mytliique, de couleur argentée, a été filmé par une équipe du Musée scienti- fique national japonais en collaboration avec la chaîne de télévision publique japonaise NHK et la chaîne spécialisée Discovery Channel. Le calamar géant, dont le nom scientifique est ar- chiteuthis, a été repci'c par 6.30 mètres de fond par une équipe en sous-marin dans le Pacifique nord. Le submersible a suivi le géant jusqu’à 900 m de profondeur avant qu ’il ne disparaisse dans les abysses. Le seul corps du calamar a été évalué à 3 m de long. Sa longueur totale a été estimée à 8 m, en l’absence de ses deux princi- paux tentaailes qui étaient sectionnés. ■
20
Novapex / Société 15(1), 10 mars 2014
Grandes marées de l’année 2014
Christiane DELONGUEVILLE et Roland SCAILEET
- Enfin une bonne année avec des marées de 1 1 5 en mars et en septembre. En février, août et octobre, les marées présenteront également de beaux coefficients supérieurs à 1 10. Voilà de quoi nous réjouir !
Coefficients (> 100) des pleines mers à Brest
(Les marées basses correspondantes sont donc particulièrement intéressantes à prospecter.)
|
Janvier |
Mercredi 1 |
(96)- 100 |
|
Jeudi 2 |
104- 107 |
|
|
Vendredi 3 |
108 - 108 |
|
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Samedi 4 |
106- 103 |
|
|
Jeudi 30 |
(95)- 101 |
|
|
Vendredi 31 |
107-111 |
|
Février |
Samedi 1 |
113 - 114 |
|
Dimanche 2 |
113 - 110 |
|
|
Lundi 3 |
106- 101 |
|
|
Vendredi 28 |
(95)- 102 |
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Mars |
Samedi 1 |
108 - 112 |
|
Dimanche 2 |
114- 115 |
|
|
Lundi 3 |
114- 112 |
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|
Mardi 4 |
108 - 102 |
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Dimanche 30 |
104-107 |
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Lundi 3 1 |
109- 109 |
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Avril |
Mardi 1 |
108 - 106 |
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Mercredi 2 |
102 -(98) |
Mai
|
Juin |
Samedi 14 |
(98) - 100 |
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Dimanche 15 |
100-99 |
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Juillet |
Dimanche 13 |
101 - 105 |
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Lundi 14 |
106-107 |
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Mardi 15 |
106 - 104 |
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|
Mercredi 16 |
101- (96) |
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Août |
Dimanche 10 |
(95)- 101 |
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Lundi 1 1 |
106- 110 |
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|
Mardi 12 |
112 - 113 |
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|
Mercredi 13 |
112- 110 |
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|
Jeudi 14 |
106- 101 |
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Septembre |
Lundi 8 |
(97)- 104 |
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Mardi 9 |
109-113 |
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Mercredi 10 |
115 - 115 |
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|
Jeudi 1 1 |
114- 111 |
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|
Vendredi 12 |
106-100 |
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Octobre |
Mardi 7 |
(96)- 102 |
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Mercredi 8 |
106- 109 |
|
|
Jeudi 9 |
111 - 111 |
|
|
Vendredi 10 |
109-106 |
|
|
Samedi 1 1 |
102 - (96) |
|
Novembre |
Jeudi 6 |
(98)- 100 |
|
Vendredi 7 |
101 - 101 |
|
|
Samedi 8 |
100 -(98) |
Décembre ^ ^
Conseils pour une marée réussie et soucieuse de l’environnement : Remettez toujours les pierres déplacées en bon ordre. Observez, photographiez et n’échantillonnez que le strict nécessaire. Soyez prudents et renseignez-vous sur les heures des marées à l’endroit où vous vous trouvez.
REFERENCE :
Annuaire des Marées pour 2014 - Ports de France - Métropole - Tome 1 - SHOM (Service Hydrographique et Océanographique de la Marine) - Paris - 256 p.
Wimereux (Pas-de-Calais)
Les données reprises dans cet article peuvent également se retrouver sur notre site Internet :
http://www.societe-belge-de-malacologie.be
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VOL 1 5 (2) |
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SOMMAIRE |
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Articles originaux - Original articles |
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B. Landau & R. Houart New Muricidae (Mollusca: Neogastropoda) from the Lower Miocene Cantaure Formation of Venezuela |
23 |
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C. Vilvens |
New species and new records of Calliostomatidae (Gastropoda: Trochoidea) from eastern and central Indo- Pacific |
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C. Delongue ville & R. Scaillet |
Y Hexaplex saharicus sahariens (Locard, 1897), distribution et illustrations des variations intra-spécifiques |
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C. Vilvens, R. Houart, E. Meuleman & M. Alexandre |
L'Assemblée Générale de la Société Belge de Malacologie du 22 mars 2014 (■ |
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(suite du sommaire en dernière page de couverture) |
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SOCIETE BELGE DE MALACOLOGIE
B. Landau & R. Hüuart
Novaphx 15(2): 23-35, K) juin 2014
New Muricidae (Mollusca: Neogastropoda) from the Lower Miocene
Cantaure Formation of Venezuela
Bernard LANDAU *
Naturalis Biodiversity Center, P.O. Box 9517, 2300 RA Leiden,
Netherlands
Centro de Geologia da Universidade de Lisboa. Campo Grande,
1749-016 Lisboa, Portugal
and
International Health Centres, Av. Infante de Henrique 7, Areias Sâo Joâo,
8200 Albufeira, Portugal bernielandau@sapo.pt
Roland HOU ART Research Associate
Institut royal des Sciences naturelles de Belgique Rue Vautier, 29, 1000 Bruxelles, Belgium. roland.houart@skynet.be
* Corresponding author
ABSTRACT, Seven new muricine records are added to that for the Lower Miocene Cantaure Formation of Venezuela, four of which are new: Siratus harzhauseri nov. sp. Ocenebra etteri nov. sp., Typhina canaliculata nov. sp. and Laevityphis jimgi nov. sp., and three are identified to genus level Phyllonoîus sp. cf. P. infrequens (Vokes, 1963), Purpurellus sp. cf. P. repetiti (Vokes, 1970) and Vitiilaria sp. cf. V. salebrosa (King & Broderip, 1832). The genus Ocenebra is recorded in the western Atlantic for the first time.
KEYWORDS. Muricidae, Mollusca, Miocene, Cantaure Formation, Venezuela, new records, new taxa.
INTRODUCTION
In his landmark monograph on the molluscan assemblage of the Cantaure Formation, Jung (1965) recorded and described ten muricine species from the deposits. Further new species were added in the 1990’s by Vokes (1992, 1994, 1995), Vermeij & Vokes (1997), and later by Vermeij (2001) and Landau & Vermeij (2010). A full list of ail species described or recorded from the Cantaure Formation subséquent to Jung’s (1965) monograph were listed by Landau & Vermeij (2010, p. l()5,appendix 1).
A short account of the geographical and stratigraphical range was given by Landau & Vermeij (2010, p. 99). This will not be repeated here, except to say that the assemblage is now considered to be late early Miocene in âge (Landau et al., 2012, p. 258, chart 18). In this contribution to the knowledge of the gastropod assemblage found in the late early Cantaure Formation
of Venezuela, we focus on new species, or taxa not previously recorded from the assemblage. This increases the number of muricine species known from the formation to 29.
Material and methods
The material described here is from the Gibson-Smith collection housed in the Naturhistorisches Muséum Basel (NHMB colL), Switzerland and the Bernard Landau collection (BL coll.), now deposited in the Naturhistorisches Muséum Wien (NHMW coll.), Vienna.
The classification adopted here is according to WoRMS (http://www.marinespecies.org/index.php). The descriptions adopt the terminology suggested by Merle ( 1999, 2001 ) (Figs 1 , 8), see next page:
23
B. Landau & R. Houar r
New Muricklae tVoni Lower Miocene of Venezuela
|
P |
Primary eord |
|
s |
secondary eord |
|
t |
tertiary eord |
|
Ad |
adapical (or adapertural) |
|
Ab |
abapical (or abapertural) |
|
SP |
Subsutural eord |
|
IP |
InfrasLitLiral primary eord (primary eord on shoulder) |
|
adis |
adapical infrasutural secondary eord (shoulder) |
|
abis |
abapical infrasutural secondary eord (shoulder) |
|
PI |
Shoulder eord |
|
P2-P6 |
Primary cords of the convex part of the teleoconch whorl |
|
sl-s6 |
secondary cords of the convex part of the teleoconch whorl |
example: si = secondary eord between PI and P2; s2 = secondary eord between P2 and P3, etc.
|
ADP |
adapertural primary eord on the siphonal canal |
|
MP |
médian primary eord on the siphonal canal |
|
ABP |
abapertural primary eord on the siphonal canal |
|
ads : |
adapertural secondary eord on the siphonal canal |
|
ms : |
médian secondary eord on the siphonal canal |
|
abs : |
abapertural secondary eord on the siphonal canal |
SYSTEMATIC PALAEON TOLOGY
Superfamily MURICOIDEA Rafinesque, 1815 Family MURICIDAE Rafinesque, 1815 Subfamily MURICINAE Rafinesque, 1815 Genus Siratus Jousseaume, 1880
Type species. - Murex senegalensis Gmelin, 1791 , by original désignation. Recent, Brazil.
Discussion. As discussed by Barco et al. (2010), Siratus Jousseaume, 1880 is doser to Vokesimurex Petuch, 1994 than it is to Chicoreus de Montfort, 1810, and ail three should be used at full generic rank. Merle et al. (2011, p. 72) discussed the difficulty in separating some species of Siratus from Vokesimurex. They suggested that well-developed secondary internai denticles and numerous columellar denticles distinguished many Siratus from Vokesimurex, but that these characters are variable. A more reliable character is the shape of the siphonal canal, which is bent in Siratus and straight in Vokesimurex. The new species from Cantaure has weak apertural dentition on both lips, which can be présent in either genus, but the siphonal canal, although incomplète, is clearly bent, placing it in the genus Siratus.
Siratus harzhaiiseri nov. sp.
Figures 1,9-1 1
Dimensions and type material. Holotype; NHMW 2()13/()476/()()10 (ex BL coll.) (Figs 1, 9-11), height 48.6 mm.
Eltymology. Named for Mathias Harzhauser ot the Naturhistorisches Muséum Wien, fricnd of the senior author, in récognition of his contributions to palaeomalacology.
Type locality. 1 km Southwest of Casa Cantaure, about 10 km west of Pueblo Nuevo, Falcon, Venezuela (= locality GSIPGNA of Gibson-Smith & Gibson-Smith, 1979).
Type stratum. Cantaure Formation (late early Miocene: Burdigalian).
Diagnosis. A medium sized Siratus species, with a relatively elongated shell, a narrowly canaliculated suture, three varices per whorl bearing short, stout spines placed mid-whorl, with three intervarical axial ridges on early whorls, four with prominent tubercles mid-whorl on last Wi whorls.
Description. Shell medium-sized, fusiform, spire tall. Protoconch not preserved. Teleoconch of six high, weakly-shoLildered whorls, with a narrow, concave, SLib-horizontal subsutural area; whorls convex below. Suture impressed, shallowly canaliculated. Three elevated varices per whorl, placed at 120°, aligned almost vertical ly, bearing a short, stout spine placed just above mid-whorl corresponding to PI, giving whorls a somewhat angular appearance. Intervarical axial sculpture consisting of three nodulose ridges on early whorls, increasing to four on abapical half of penultimate whorl, where the nodules mid-whorl become more prominent. Last whorl fusoid; subsutural area with IP, adis and abis of almost equal strength. Below shoulder P2-P6 narrow, s2-s6 developed in the interspaces between the primary cords. ADP and MP developed on base, ads and ms weaker. Tertiary cords developed throughout between primary and secondary éléments. Siphonal canal below ms missing. Aperture of moderate size, ovate. Outer lip convex, margin erect bearing 16 denticles at the inner edge. Labral varix moderately expanded, roLinded, bearing straight spine at shoulder. Anal sinus broad, shallow; siphonal canal incomplète, but long.
24
B. Landau & R. Hou art
NOVAPBX 15(2): 23-35, 10 juin 2014
narrow, open, pointing slightly to left. Cokimellar lip adapically, erect abapically, beariiig elongatcd
well dclimited, narrovvly expandcd, adhèrent denlieles along its entire length.
Figure 1. Siratiis harzhauseri nov. sp., holotype NHMW 2013/0476/0010 (ex BL eoll.), height 48.6 mm. 1 km Southwest of Casa Cantaure, about 10 km west of Pueblo Nuevo, Falcon, Venezuela, Cantaure Formation, Burdigalian, late early Mioeene.
Discussion, Siratiis harzhauseri nov. sp. is most similar to the Caribbean Plioeene-Reeent S. formosus (Sowerby, 1841), but differs in having a more elongated fusiform shell shape, in having a shailowly eanalieulated suture, in having PI and henee the shoulder spine plaeed lower, eloser to mid-whorl and shorter, and in having the intervarieal axial ridges more strongly nodulose mid-whorl on the last l'A whorls. Siratus domingensis (Sowerby, G. B. I, 1850) from the early Plioeene Gurabo Formation of the Dominiean Republie is similar to S. formosus and was eonsidered to be aneestral to it by Vokes (1989). Apart for a few exeeptionally large speeimens, S. domingensis is smaller than either S. formosus or S. harzhauseri, it has smaller spines and a more reeurved siphonal canal. Siratus domingensis has three-five intervarieal axial ridges, which are narrower and the suture is not eanalieulated. Siratus articulatus (Reeve, 1845) from the Plioeene-Reeent Caribbean can easily be separated as PI, P3 , P5 and P6 form more or less long spines at the apertural varix and the suture is not eanalieulated. Another closely similar species is the early Pleistocene to Recent Caribbean Siratus springeri (Bullis, 1964), but this species differs from S. harzhauseri in having convex rather than angular whorl profile, in having only two or three axial intervarieal ridges, in having the labral varix deeply excavated on the abapertural side, and in most speeimens PI , P5, P6 are strongly developed.
Two further Siratus species occur in the Cantaure assemblage: S. quirosensis (F. Hodson, 1931), which is much smaller than S. harzhauseri, with only two axial intervarieal ridges that are more prominent than
in S. harzhauseri, and PI, P3 and P5 are strongly developed. The second, Siratus denegatus (Jung, 1966), is quite different from al) its congeners in having a rather triangular shell, a sharply carinate last whorl, only one axial intervarieal ridge, very fine spiral sculpture, and PI developed into a short, sharp spine at the shoulder.
Genus Phyllonotiis Swainson, 1 833
Type species. - Murex imperialis (var. a) Swainson, 1833, by subséquent désignation, Swainson, 1833, pl. 109 (= Murex imperialis Swainson, 1831) (not Fischer de Waldheim, 1807) (= Murex margaritensis Abbott. 1958). Recent, West Atlantic.
Phyllonotus sp. cf. P. infreq liens (Vokes, 1963) Figures 12-13
cf. 1963. Murex {Phyllonotus) infrequens Vokes. p. 1 56, pl. 1 , fig. 4.
Material, 1 specimen NHMW 2013/0476/0011 (ex BL eoll.), height 35.3 mm (Figs 12-13).
Locality. 1 km Southwest of Casa Cantaure, about 10 km west of Pueblo Nuevo. Falcon. Venezuela, Cantaure Formation, Burdigalian, late early Mioeene.
Discussion. This specimen resembles the species illustrated by Vokes (1989, 1990) as Phyllonotus infrequens from the Early Mioeene Baitoa Formation of the Dominical! Republic. However Phyllonotus
25
B. Landau & R. Houari
New Muricidae from Lowcr Miocène of Venezuela
iiifrcciiieiis from the Early Miocene Chipola Formation of Florida has a more globose shell with a less impressed suture, a more rounded aperture and less conspicLiOLis secondary and tertiary spiral cords. The specimen from the Baitoa Formation and our specimen from the Cantaure Formation are somewhat related. although the Cantaure one is more globose, has a more impressed suture and a narrower siphonal canal .
Genus Piirpiirelliis Jousseaume, 1880
Type species. - Murex gambiensis Reeve, 1845, by original désignation. Recent, West Africa.
Emerson & D’Attilio (1969) demonstrated that, on the basis of radular morphology, Purpiirelliis Jousseaume, 1880 was closely related to Pterynotus. Purpiirelliis differs from Pterynotus in having a sealed siphonal canal and an operculum with a central nucléus, instead of apical in Pterynotus s.s. The use of the taxon at full generic rank is now generally accepted (Landau et al., 2007, Merle et al., 2011, Bouchet & Houart, 2013).
Piirpurellus sp. cf. P. repetiti (Vokes, 1970) Figures 14-17
cf. 1970 Pterynotus (Piirpurellus) repetiti Vokes, p. 16, pl. 3, fig. 4.
Material. 1 incomplète specimen, NHMW 2013/0476/0008 (ex BL coll.), height 27.2 mm (Figs 14-15); 1 incomplète specimen, NHMW
2013/0476/0009 (ex BL coll.), height 32.4 mm (Figs 16-17).
Locality. 1 km Southwest of Casa Cantaure, about 10 km West of Pueblo Nuevo, Falcon, Venezuela (=locality GSIPGNA of Gibson-Smith & Gibson- Smith, 1979), Cantaure Formation (early Miocene: Burdigalian).
Discussion. Vokes (1989, p. 55) commented that Piirpurellus species could be difficult to separate. The two incomplète specimens illustrated here resemble the specimen described as Piirpurellus repetiti (Vokes, 1970) from the “Silverdale Beds” of North Carolina, originally considered Early Miocene, but placed in the Late Oligocène Chickasawhayan Stage by Gibson (1977, p. 202; 1983, p. 38). This species is characterised by its relatively tall spire, moderately expanded varices, with short straight spines at the shoLilder, single sharp intervarical node, and very weak spiral sculpture. It is most like the European Miocene-Pliocene Purpurellus veranyi (Paulucci, 1866) (.see Landau et al., 2007; Merle et al., 2011), but this species has broader intervarical nodes and a relatively smaller, more circular aperture. Apart from the.se constant différences, most specimens of P . veranyi are broader, hâve stronger, recurved spines at the shoLilder, slightiy more prominent spiral sculpture.
and more expanded labral varix. However, given the badly preserved material from the Cantaure Formation and the few specimens involved, the identification remains doubtful.
The only other western Atlantic congener is Purpurellus inirificiis Vokes, 1989 from the early Pliocène Gurabo Formation of the Dominican Republic, which is quite different from P. repetiti and P. veranyi in having greatly expanded wing-like varices, similar to the Recent West African P. gambiensis (Reeve, 1845).
In Europe the fossil record also starts in the Late Oligocène with an undescribed species listed by Lozouet (1986, p. 377) from the Atlantic Late Oligocène of France. Today Purpurellus has a disjunct distribution, P. gambiensis (Reeve, 1845) from West Africa and two species, P. pinniger (Broderip, 1833) and P. macleani (Emerson & D’Attilio, 1969) from the Pacific coast of Tropical America. In the fossil record, specimens of Purpurellus are alvvays rare, and although the first record is Late Oligocène in both the eastern and western Atlantic, it is likely that the genus is a Palaeogene offshoot of Pterynotus, which originated in the Tethys, immigrated to the western Atlantic and colonised the Tropical American Pacific before the closure of the central American Seaway, sLibsequently dying out in the western Atlantic.
Subfamily OCENEBRINAE Cossmann, 1903 Genus Ocenebra Gray, 1847
Type species. - Murex erinaceus Linnaeus, 1758, by monotypy. Recent, northeastern Atlantic, Mediterranean.
Ocenebra etteri nov. sp.
Figures 18-23
Dimensions and type material. Holotype, NHMW 2013/0476/0018 (Figs 20-21), height 22.3 mm; paratype 1, NHMW 2013/0476/0017 (Figs 18-19). height 17.5 mm; paratype 2, 2013/0476/0019, height 22.4 mm; paratype 3, NHMW 2013/0476/0021, height 12.1 mm.
Other material. NHMW 2013/0476/0020 (1), height 14.0 mm, juvénile (Figs 22-23).
Etymology. Named after Dr. Walter Etter of the Naturhistorisches Muséum Basel, Switzerland, in récognition of his help and support.
Type locality. 1 km Southwest of Casa Cantaure, about 10 km west of Pueblo Nuevo, Falcon. Venezuela (= locality GSIPGNA of Gibson-Smith & Gibson-Smith, 1979).
Type stratum. Cantaure Formation (late early Miocene; Burdigalian).
26
B. Landau & R. Hou art
NovAPHX 15(2): 23-35. 10 juin 2014
Diagnosis. A medium sized Ocenehm specics with a strongly nodulose shouldcr, Uiree varices per vvhorl on the last three whorls, with one intervarical node, spiral eords P1-P5 equal in slrength, P6 slightiy narrovver, with a single interealated secondary eord, aperture small with very shallow ID and strong D1-D5 within and a short, sealed siphonal canal.
Description. Shell medium sized, heavy, nodose. SubsLitural ramp broad, weakly sloping, slightiy eoneave. Spire high with teleoconch up to five or six strongly shouldered, nodose whorls. Suture adpressed. Protoconeh unknown. Axial sculpture of teleoconch whorls consisting of high ribs and high, broad, varices. Eaeh varix with short, blunt nodes extending from primary spiral cords. Antepenultimate, penultimate and last whorls with three varices and a single, strong, prominent, intervarical node. Other spire whorls with narrow axial ribs. Last whori broad. SubsLitural area with adis, IP, abis, followed on convex part of vvhorl by PI , si , P2, s2, P3, s3, P4, s4, P5, s5, P6, s6; ADP and MP on siphonal canal. P1-P6 almost similar in size and strength, P6 weakly narrovver. Secondary cords obviousiy narrower than primary cords. Siphonal canal belovv MP missing or damaged. Aperture moderately small, ovate. Columellar lip narrow, smooth, adhèrent. Anal notch shallow, broad. Outer apertural lip with very shallow ID and strong D1-D5 within. Labral varix rounded, strong, broad, with small nodes. Siphonal canal short, narrow, ventrally sealed, partly broken.
Discussion. VermeiJ & Vokes (1997, p. 72) restrict the genus Ocenebra to "a relatively small number of Miocene to Recent species from western Europe, the Mediterranean région and Tropical West Africa. These are characterized by the tendency to form three varices on the last whori, by the presence of six to eight primary spiral cords on the last whori, a crenulated outer lip without a labral tooth, an adhèrent or very lightly erect inner lip, and six weak to strong denticles on the inner side of the outer lip. In species with varices, the latter are separated from eaeh other by a single intervarical node".
The Tropical West African species are now classified in other généra. However, Houart & Sirenko (2003) also assigned three Recent and fossil species from the northwestern Pacific to Ocenebra.
Two more or less related species were assigned to Miocenebra: M. silverdalensis (Vokes, 1963) and to Fenoligniim: F. umbiFicatum VermeiJ & Vokes, 1997, by VermeiJ & Vokes (1997), both species from the Belgrade Eormation, North Carolina.
This is the first occurrence of the genus Ocenebra in the tropical western Atlantic and thercfore this new species cannot be compared to any Caribbean fossil or Recent taxon. Ocenebra etteri nov. sp. is far more reminiscent of some of the Early Miocene Aquitanian and Burdigalian species from the French Aquitaine Basin (see Lozouet et al., 2001 , pl 22, 23).
Genus VitiiUma Swainson, 1840
Type species. — Murex niiliaris Gmelin, 1791, by monotypy. Recent, western Indo-Pacific.
Note. - The placement of this genus in the Ocenebrinae Cossmann, 1903 is provisional, as molecular data prcsented by Barco et al. (2010) showed the subfamily to be polyphyletic.
Vitularia ,sp. cf. V. salebrosa (King & Broderip, 1832)
Figures 24-27
cf. 1832 Murex salebrosa King & Broderip, p. 347.
Material. 1 incomplète specimen, NHMW 2012/0197/0008 (ex BL colL), height 29.1 mm (Figs 24-25); 1 incomplète specimen, NMB H20169, NMB locality 12842, height 72.2 mm (Figs 26-27).
Locality. 1 km Southwest of Casa Cantaure, about 10 km west of Pueblo Nuevo, Falcôn, Venezuela ( = locality GSIPGNA of Gibson-Smith & Gibson-Smith, 1979), Cantaure Formation (early Miocene: Burdigalian).
Discussion. A Vitularia species occurs in the Cantaure formation, which is difficult to characterise as only two incomplète specimens are available to us, and they are somewhat different from eaeh other. The first of these is smaller and probably represents a subadult stage (Figs 24-25). The shell is slender and the spire high. The carina on the spire whorls is sharp and runs mid-whorl. The subsutural ramp is broad and weakly concave, below the carina the vvhorl is straight-sided and tapers inwards to the suture. The axial sculpture consists of eight prosocline ribs developed into prominent rounded tubercles at the shoulder. The ribs cross the subsutural ramp as narrow lamellae. The surface is pustulose, a feature of the genus. Unfortunately the adapical portion of the last vvhorl and aperture are missing. This is the specimen discussed by Landau et al. (2013). The second specimen (Figs 26-27) is larger, but strongly abraded and incomplète. The spire is also high. and the shoulder bears rounded tubercles. However. it differs in the position of the carina on the spire whorls, which is doser to the adapical suture, giving the spire a conical rather than scalate shape. The two Vitularia .specimens from Cantaure probably represent a single species, as these différences in the position of the suture and spire shape can also be seen in the European Miocene Vitularia linguabovis (Basterot, 1825) (Landau et al., 2013).
The genus is not speciose. In the Caribbean faunas only two species are known. Vitularia clominicana Vokes, 1989 from the early Pliocène Gurabo Formation of the Dominican Republic is lovv spired, and according the illustrated holotype, has large round
27
B. Landau & R. Houarf
New Muricidae from Lower Miocene of Venezuela
tubercles at the shoulder, which is placed high on the last whorl. At the time of description only three specimens were available to Emily Vokes, with a maximum height of 28.2 mm. Subséquent collections made by the senior author hâve brought further specimens to light, which attain a greater maximum height of 53.9 mm. As the shell increases in size the penultimate and last whorls become rounded and lose
the tubercles at the shoulder. In the largest specimens from the Dominican Republic the shoulder of the last whorl is rounded and quite smooth. In view of these important and, so far undocumented ontogénie changes, we take the opportunity to illustrate further specimens from the Dominican Republic (Figures 2- 7).
Figures 2-7. Vitiilaria dominicana Vokes, 1989, Tulane locality 1215 (see Vokes, 1989), Gurabo River, Gurabo Formation, Lower Pliocène, Dominican Republic. Figs 2-3 NHMW 2013/0476/0005, height 40.8 mm; figs 4-5 NHMW 2013/0476/0006, height 53.7 mm; figs 6-7 NHMW 2013/0476/0007, height 49.5 mm.
Vokes (1989) compared the Dominican shell to the Recent Indo-Pacific species Vitularia miliaris (Gmelin, 1791), but large specimens of this species always hâve large tubercles at the shoulder, which remains angular and placed high on the last whorl.
The second species is Vitularia salebrosa (King & Broderip, 1832) (= V. ecuadoriana Marks, 1951 = V. lingiiahison Vokes, 1967). This species occurs in the Caribbean Neogene in the late Pliocène Agueguexquite Formation of Mexico and Pinecrest Formation of Florida, in the Panamic faunas from the early Pliocène Esmeraldas Group of Ecuador, through the Pleistocene of Baja California, Ecuador and the Galapagos, and is found today from Baja California to Peru. It is one of the paciphile species discussed by Landau et al. (2009). Vitularia salebrosa is tall-spired, like the Cantaure specimens, but seems to hâve smaller and more numerous tubercles at the shoulder. Vitularia linguabovis (Basterot, 1825), which occurs in the early-late European Miocene, is broader, usually with a less elevated spire, and more numerous tubercles at the shoulder. We wait in the hope ot finding better specimens from Cantaure, which is the earliest occurrence of the genus in the tropical American Neogene assemblages.
Subfamily TYPHINAE Cossmann, 1903 Genus Typhina Jousseaume, 1880
Type species. — Typhis belcheri Broderip, 1833 (= Murex cleryi Petit, 1840) by original désignation. Recent, West Africa and Brazil.
= Tr/Z/Yv/?/? A Jousseaume, 1882
Talityphis Jousseaume, 1882 and Typhina Jousseaume, 1880 were considered congeneric by Houart (2002).
Typhina canaliculata nov. sp.
Figures 8, 28-34
Dimensions and type material. Holotype; NHMW 2013/0476/0001 (ex BL coll.) (Figs 28-31), height 28.6 mm; paratype 1; NHMW 2013/0476/0002 (ex BL coll.) (Figs 8, 32-34), height 23.8 mm; paratype 2; NHMW 2013/0476/0003 (ex BL coll.), height 28.9 mm. paratype 3 NMB H20I7I (NMB locality 17520), height 19.9 mm; paratype 4 NMB H20I70 (NMB locality 17516), height 27.9 mm; paratype 5 NMB H 19068 (NMB locality 1 75 16), height 27.7 mm.
28
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NOVARHX 15(2): 23-35. 10 juin 2014
Other material. Maximum height 30.3 mm; 10 specimens NHMW 2013/0476/0004 (ex BL coll.), NMB locality 17516 NMB coll. (6), same locality.
Etymology, Name rel'lecting the deeply canaliculated suture seeu iu this species.
Type locality. 1 km Southwest of Casa Cautaure, about 10 km west of Pueblo Nuevo, Falcou, Venezuela (= locality GSIPGNA of Gibsou-Smith & Gibsou-Smith, 1979).
Type stratum. Cautaure Formation (late early Mioceue; Burdigaliau).
Diagnosis. A medium-sized Typhina species, with a relatively high spire, strougly concave subsutural area resultiug iu a deeply canaliculated suture, four varices per whorl, short tubes, five weakly developed spiral cords formiug weakly recurved spiues on the varices, and a moderately expanded labral varix.
Description. Shell medium-sized, stout; spire
relatively high. Protocouch uot preserved. Six or scveu sharply-augular, straight-sided teleocouch whorls, with uarrow, deeply concave subsutural area. Suture impresscd, deeply canaliculated. Four varices per whorl. Tubes (PI) of moderate leugth, poiutiug laterally and abaperturally, placed doser to precediug thau SLicceediug varices. A lamiuar partition above the apcrture Crossing the shoulder joins the varix to the varix of the preceding whorl. Below tube shell surface very weakly swollen and two Unes pass anteriorly, one being the margin of the old mouth, the other, in advance of the tube, being similar in character and indicating another arrest in growth. P2-P6 weakly developed, forming small, posteriorly-reflected spiues over the varices. Surface bearing faint growth lines. Aperture small, ovate, bordered by raised rim; labral varix constant in width, moderately expanded, consisting of inner band bearing spiral cords and outer flange with a scalloped edge formed by spines at termination of P2-P6. Siphonal canal closed, broad, relatively short, pointing to the right and abaperturally.
Anal tube = P1
Partition
Inner band p5 Outer flange Siphonal canal
Figure 8. Typhina canaliculata nov. sp., paratype I NHMW 2013/0476/0002 (ex BL coll.), height 23.8 mm, 1 km Southwest of Casa Cantaure, about 10 km west of Pueblo Nuevo, Falcôn, Venezuela. Cautaure Formation, Burdigaliau, late early Mioceue.
Discussion. The four varices and tubes per whorl, the tubes being placed doser to the preceding than SLicceeding varix, the apertural varix greatly expanded and the partition above the aperture place this new species in the genus Typhina Jousseaume, 1880. Typhina canaliculata nov. sp. resembles the Recent and type species T. helcheri (= T. cieryi) but it differs in having a comparatively larger shell with a higher spire, a straighter, less upward bent and shorter P2 spine instead of long and strongly adapically recurved
in T. helcheri, a more sculptured. more strongly folded apertural varix and a comparatively broader siphonal canal.
Typhina canaliculata nov. sp. is also closely similar to Typhina ohesa (Gabb. 1873), which is widespread in the Caribbean early Mioceue (Vokes, 1989), but differs in being less broad and in having a taller spire, whereas T. ohesa lias a very short, depressed spire. The subsutural area in T. ohesa is broad and weakly concave, whereas in T. canaliculata it is narrower and
29
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New Miiricidae tVom Lower Miocene of Venezuela
strongly coneave. resiilting in a deepiy canaliculated suture, bordered by a high-ridged shoulder. In Typhina caïuiliciilala the spiral cords are more strongly developed than in T. obesa, forming short recLirved spines at the varices on the last whorl. In T. ohesa the spiral sculpture is very weakly developed, hardly marked over the sharp, straight varices. Lastly, the aperture is relatively smaller and the labral varix narrower than in T. ohesa. Typhina e.xpansa (Sowerby, 1874) from the early Pliocène to Recent Caribbean, is very closely similar to T. ohesa (différences between the two were discussed by Vokes, 1989, p. 332), but this species can also be distinguished from T. eanaliciilata in having a broader shell shape and in having a wider apertural varix, but above ail by the less concave subsutural area. Typhina alata (Sowerby, 1850) from the early Pliocène Gurabo Formation of the Dominican Republic and Pliocène formations of Ecuador differs in having a more fusiform shell shape, a less depressed subsutural area, a wider labral varix and in forming an intervarical node below the tubes, a character not seen in T. ohesa, T. e.xpansa or T. canaliculata.
The earliest member of the genus in the Caribbean was thought to be Typhina precursor (Keen & Campbell, 1964) from Las Perdices Shale of Colombia, which, according to Gertman (1969), was dated as Aquitanian early Miocene. This species also occLirs in the Manzanilla Formation of Trinidad, dated by Gertman as early Miocene. The âge of the Las Pedices Group is now considered middle-late Miocene (Landau et al., 2012, chart 2), and the Manzanilla Formation is considered to be late Miocene (Landau et al., 2012, chart 15). Typhina preeitrsor is a very large solid species, 47-49 mm in height, which also seems to hâve a narrow shoulder, but differs importantly from T. canaliculata and other Typhina species in having no spiral sculpture developed.
Other Caribbean congeners are: Typhina pterina (Gardner, 1947) from the middle Miocene Shoal River Formation of Florida and T. eiicteana (Woodring, 1970) from the late Miocene Gatun formation of Panama, which are both much smaller, with a more fusiform shell shape. The subsutural area in both of
these is concave, but not as deep as in T. canaliculata. Typhina eucteana is further distinguished by its long, narrow siphonal canal. Typhina siphon (Woodring, 1928) from the Pliocène Bowden Formation of Jamaica is a more slender, fusiform species, with recLirved spines at the shoulder, and a longer siphonal canal. Typhina acuticosta (Conrad, 1830) from the middle Miocene St. Mary Formation of Maryland and late Miocene Choctawhatchee Formation of Florida is quite different, with a slender shell shape and acutely angled, straight-sided whorls. Typhina cannenae (Gertman, 1969) from the late Pliocène Agueguexquite Formation of Mexico has a broader shell shape, more prominent spiral sculpture and a far more expanded apertural varix than T. canaliculata. The Recent Panamic Typhina latipennis (Dali, 1919) has weaker spiral sculpture than the fossil Cantaure species, and a greatly expanded flange on the apertural varix.
Genus Laevityphis Cossmann, 1903
Type species. - Typhis coronarius Deshayes, 1865 {Laevityphis muticiis (Sowerby, 1835)], by original désignation. Eocene, France.
Laevityphis jiingi nov. sp.
Figures 35-38
Typhis (Laevityphis) sawkinsi. Jung, 1965, p. 525, pl. 70, figs 7, 8 [not Laevityphis sawkinsi (Mansfield 1925)].
Dimensions and type material. Holotype; NHMW 2013/0476/0012 (ex BL coll.) (Figs 35-36), height 28.3 mm; paratype 1; NHMW 2013/0476/0013 (ex BL coll.) (Figs 37-38), height 27.0 mm; paratype 2; NHMW 20 13/0476/00 14^ (ex BL coll.). height 27.1 mm; paratype 3; NHMW 2013/0476/0015 (ex BL coll.), height 23.8 mm; paratype 4 NMB H18542, height 23.0 mm; paratype 5 NMB H 18543, height 20.7 mm; paratype 6 NMB H 137 19, height 23.0 mm [specimen figured by Jung, 1965, pl. 70, figs 7, 8 as Typhis (Laevityphis) sawkinsi Mansfield].
Figures 9-23
9-1 1. 57/Ï///7.V harzhauseri nov. sp., holotype NHMW 2013/0476/0010 (ex BL coll.), height 48.6 mm.
12-13. Phyllonotus sp. cf. P. infrecpiens (Vokes, 1963), NHMW 2013/0476/001 1 (ex BL coll.), height 35.3 mm. 14-17. Purpurellus sp. cf. P. repetiti (Vokes, 1970)
14- 15. NHMW 2013/0476/0008 (ex BL coll.), height 27.2 mm; 16-17. NHMW 2013/0476/0009 (ex BL coll.), height 32.4 mm.
15- 23. Ocenehra etteri nov. sp.
18-19. Paratype 1 , NHMW 2013/0476/0017, height 17.5 mm; 20-21. Holotype, NHMW 2013/0476/0018, height 22.3 mm; 22-23. Juvénile, NHMW 2013/0476/0(^20, height 14.0 mm.
Ail: I km Southwest ol Casa Cantaure, about 10 km west ot Pueblo Nuevo, Falcôn, Venezuela, Cantauie Formation, Burdigalian, late early Miocene.
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New Muricidae from Lower Miocene of Venezuela
Other material. Maximum height 31.7 mm; 20 specimens NHMW 2013/0476/0016 (ex BL coll.), NMB locality 17516 NMB coll. (66), NMB locality 1 7520 NMB ( 100+), same locality.
Etymology. Named in honour or Peter Jung, in récognition of his work on the Cantaure assemblage.
Type locality. 1 km Southwest of Casa Cantaure, about 10 km west of Pueblo Nuevo, Falcôn, Venezuela (= locality GSIPGNA of Gibson-Smith & Gibson-Smith. 1979).
Type stratum. Cantaure Formation (late early Miocene: Burdigalian).
Diagnosis. A medium-sized Laevityphis species, with a high spire, a narrow, concave subsutural area, four varices per whorl placed in the middle of the intervarical area, bearing spines pointing adapically and inwards, short tubes in the intervarical spaces, an almost smooth surface, a small aperture, a naiTowly expanded labral varix, and a long, narrow siphonal canal.
Description. Shell medium-sized, elongate; spire high. Protoconch paucispiral, consisting of two elevated, bulbous whorls. Six or seven sharply-angular teleoconch whorls, with narrow, concave, sub- horizontal subsutural area, weakly convex below shoulder to suture. Suture impressed, undulating. Four simple, convex varices per whorl, with a large, sharp spine at the shoulder of each varix pointing adapically and inwards; end usually broken away and appearing as small tube-like structures, but not connecting to interior of shell. Tubes (PI) of moderate length, separated from varices, pointing laterally and abaperturally, placed in the middle of the intervarical area. A ridge running abapically from the tube dividing the intervarical area into two almost equal halves. Last whorl relatively short, convex, strongly constricted at the base. P2-P6 not clearly developed.
very weak spiral sculpture just visible on penultimate and last whorls. Aperture small, ovate, bordered by raised rim; labral varix constant in width, moderately expanded. Siphonal canal closed, narrow, long, pointing to the right and abaperturally at distal end.
Discussion. The four varices and tubes per whorl, the tubes not being attached to the varix, the varix bearing a spine pointing apically and the narrowly expanded apertural varix place this new species in the genus Laevityphis Cossmann, 1903 (Gertman, 1969). In this genus the tube is usually placed doser to the succeeding than the preceding varix, although in Laevityphis jitngi nov. sp. the tube is placed almost midway between the varices.
This species was first identified by Jung (1965) as Typhis (Laevityphis) sawkinsi (Mansfield, 1925) from the early-middle Miocene Brasso Formation of Trinidad, although the author noted that the Cantaure shells were not identical with those from Trinidad. When compared to the holotype figured by Gertman (1969, pl. 7, fig. 1), the Trinidadian species is much stockier, with a far shorter spire, the varices are far wider and more prominent than in L.jiingi, the base is less strongly constricted and the siphonal canal is broader and shorter than in the Venezuelan species. The fact that the Venezuelan and Trinidadian species were not conspecific was noted by Vokes (1989, p. 81), who considered the Cantaure shells to be a rather large variant of Laevityphis linguiferus (Dali, 1890) a species common in the early Miocene Chipola Formation of Florida. Indeed, Vokes (1989) considered the two identical, apart from their size. We hâve compared the Cantaure material with numerous specimens of L. linguiferus from the Chipola Formation and cannot agréé with this conclusion. Laevityphis jungi is indeed a much larger shelled species, adults ranging from 22.7 mm - 31.7 mm in height, whereas the largest specimens of L. linguiferus do not attain 15 mm in height. Further différences can be seen in the spire, which is much taller in the Venezuelan species and the last whorl, which is more
Figures 24-38
24-27. Vitularia sp. cf. V. salebrosa (King & Broderip, 1832)
24-25. NHMW 2012/0197/0008 (ex BL coll.), height 29.1 mm; 26-27. NMB H 12842, height 72.2 mm. 28-34. Typhina canaliculata nov. sp.
28-31. Holotype; NHMW 2013/0476/0001 (ex BL coll.), height 28.6 mm; 32-34. Paratype 2; NHMW 2013/0476/0003 (ex BL coll.), height 28.9 mm.
35-38. Laevityphis jungi nov. sp.
35-36. Holotype; NHMW 2013/0476/0012 (ex BL coll.), height 28.3 mm; 37-38. Paratype 1; NHMW 2013/0476/0013, (ex BLcoll.), height 27.0 mm.
Ail: I km southwe.st of Casa Cantaure, about 10 km west of Pueblo Nuevo, Falcôn, Venezuela, Cantaure Formation, Burdigalian, late early Miocene.
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New Muricidae from Lower Miocene of Venezuela
eonstrieted at the base. The proximal end of the siphonal canal is proportionally broader in L. lingiiifenis than in L. jiingi, but then narrows rapidiy at the distal end to form a very small siphonal exit, whereas in L. Jiingi the narrowing in graduai and the siphonal exit proportionally wider. The sculpture, or lack of it, is similar in both species.
Several congeners occur in the tropical American fossil record. The oldest member of the group, L. gracilis (Conrad. 1833) from the late middle Eocene Gosport Formation of Alabama, USA also has a slender shell and tall spire, but is distinguished by its crenulated varices. The other Miocene and Pliocène congeners, such as L. costaricensis (Olsson, 1922), L. hiillisi (German, 1969), L. apheles (Vokes, 1989) and L. spinirectiis (Vokes, 1989) ail differ in having smaller, less slender shells with lower spires.
Acknowledgements
We would like to thank Emily Vokes for her comments and for acting as referee. Also to Walter Etter and Olivier Schmidt of the Naturhistorisches Muséum Basel, Switzerland, for access to the PPP collection.
REFERENCES
Barco, A., Claremont, M., Reid, D.G., Houart, R., Bouchet, P., Williams, S.T., Cruaud, C., Couloux, A., Oliverio, M. 2010. A molecular phylogenetic framework for the Muricidae, a diverse family of carnivoroLis gastropods. Molecular Phylogenetics and Evolution 56: 1025-1039.
Bouchet, P. & Houart, R. 2013. World Register of Marine Species.
http://www.marinespecies.org/aphia.php?p=taxdet
ails&id=225362. Accessed on 19 November 2013. Emerson, W.K., D’Attilio, A. 1969. Remarks on the taxonomie placement of Purpurellus Jousseaume, 1880, with the description of a new species. The Veliger 12: 145-148.
Gertman, R.L. 1969. Cenozoic Typhinae (Mollusca: Gastropoda) of the western Atlantic région. Tulane Studies in Geology 7: 143-191 .
Gibson-Smith, J. & Gibson-Smith, W. (1979). The genus, Arcinella (Mollusca: Bivalvia) in Venezuela and some associated faunas. Geos 24: 1 1-32. Gibson, T.G. 1 911 . In Géologie and hydrologie principles, processes, and techniques; Géologie Survey Research 1977. United States Geological Survey Professional paper 1050: 153-221.
Gibson, T.G. 1983. Stratigraphy of Miocene through lower Pleistocene strata of the United States central Atlantic Coastal Plain. In Ray, C:E., ed., Geology and paleontology of the Lee Creek mine, North Carolina, 1. Smithsonian Cotrihutions to Paleohiology 53: 35-80.
Houart, R. 2002. Description of a new typhine (Gastropoda: Muricidae) from New Caledonia
with comments on some generic classifications within the SLibfamily. Venus S\ (3-4): 147-159.
Houart, R. & Sirenko, B. 2003. Review of the Recent species of Ocenehra Gray, 1847 and Ocinehrellus Jousseaume, 1880 in the Northwestern Pacific. Ruthenica 13 (1): 53-74.
Jung, P. 1965. Miocene Mollusca from the Paraguana PeninsLila, Venezuela. Bulletins of American Paleontology 49 (223): 387-644.
King, P.P. and Broderip, W.J. 1832. Description of the Cirrhipeda, Conchifera and Mollusca, in a collection formed by the Officers of H.M.S. Adventure and Beagle employed between the years 1826 and 1830 in survey ing the Southern Coasts of South America, including the Straits of Magalhaens and the Coast of Tierra del Fuego. Zoological Journal 5: 332-349.
Landau, B.M., Harzhauser M., Islamoglu, Y., Silva, C.M. da 2013. Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin, Turkey. Cainozoic Research 1 1-13: 3-584.
Landau, B.M., Houart, R., Silva, C.M. da 2007. The early Pliocène Gastropoda (Mollusca) of Estepona, Southern Spain, 7. Muricidae. Palaeontos 1 1: 1-87.
Landau, B., Petit, R.E., Etter, W., Silva, C.M. da 2012. New species and records of Cancellariinae (Caenogastropoda) from tropical America, together with a catalogue of Neogene to Recent species from this région. Cainozoic Research 9: 193-279.
Eandau B.M. & Vermeij, G.J. 2010. A new species of Plicopurpura (Mollusca: Rapaninae) from the Lower Miocene Cantaure Formation of Venezuela. Novapex 1 1: 99-106.
Landau B.M., Vermeij, G.J., Silva, C.M. 2009. Pacific éléments in the Caribbean Neogene gastropod fauna: the source-sink model, larval development, disappearance, and faunal units. Bulletin de la Société Géologicjue de France 180(4): 249-258.
Lozouet, P. 1986. Les Gastéropodes prosobranches de l’Oligocène supérieur du Bassin de l’Adour (systématique, paléoenvironnements, paléoclimatologie, paléobiogéographie). Diplôme de l’E.P.H.E., 475pp.
Lozouet, P., Lesport, J.F., & Renard, P. 2001 .
Révision des Gastropoda (Mollusca) du stratotype de l’Aquitanien (Miocène inf.): site de Saucats ‘Lariey’, Gironde, France. Cossnianniana (hors série 3): 1-189.
Merle, D. 1999. La radiation des Muricidae
{Gastropoda : Neogastropoda) au Paléogène: approche phylogénéticpie et évolutive. Paris. Unpublished thesis. Muséum national d'Histoire naturelle : i-vi,499 pp.
Merle, D. 2001 . The spiral cords and the internai denticles of the outer lip in the Muricidae: terminology and methodological comments. Novapex, 2 (3), 69-7 1 .
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Merle, D., Garrigues, B., Pointier, J.P. 2011. Fossil and Recent Miiricidae ofthe World. Part Mnrieinae. Haekenheim (ConehBooks): 648 pp.
Swainson, W. 1820-33. The Zoologieal illustrations, or original figures and deseriptions of new, rare, or interesting animais, seleeted ehiefly front the elasses of ornithology, entomology, and eonehology. London, ser. I , vols 1-3, pis 1-182 ( 1 820-23); ser. 2, vols 1-3, pis 1-136(1829-33).
Vermeij G.J. 2001 . Distribution, history, and taxonomy ofthe Thaïs elade (Gastropoda: Murieidae) in the Neogene of Iropieal America. Journal of Paleontolo^y 75: 697-705.
Vermeij G.J. & Vokes E.H. 1997. Cenozoic Murieidae ofthe western Atlantic région. Part 12. The subfamily Ocenebrinae (in part). Tiilane Stiidies in Geology and Paleontology 29: 69-1 18.
Vokes, E.H. 1963. Cenozoic Murieidae ofthe western Atlantic région. Part I - Mitre.x sensu stricto. Tiilane Stndies in Geology 1 : 93-123.
Vokes, E.H. 1970. Cenozoic Murieidae ofthe western Atlantic région. Part V. Pterynotiis and Poirieria. Tiilane Stiidies in Geology 8: 1-50.
Vokes, E.H. 1989. Neogene paleontology in the northern Dominican Republic, 8. The family Murieidae (Mollusca: Gastropoda). Bulletins of American Paleontology 97 (332): 5-94.
Vokes, E.H. 1990. Cenozoic Murieidae ofthe Western Atlantic région. Part VIII - Murex s. s.,
Haustelluni, Clueoreus, and Hexaplex\ additions and corrections. Tulane Stiidies in Geology and Paleontology 23\ 1-96.
Vokes E.H. 1992. Cenozoic Murieidae ofthe western Atlantic région. Part 9. Pterynotus, Poirieria, Aspella, Dernionnirex, Calotroplion, Acantliolahia , and Attiliosa', additions and corrections. Tulane Stiidies in Geology and Paleontology 25: 1-108.
Vokes E.H. 1994. Cenozoic Murieidae of the western Atlantic région. Part 10. The subfamily Muricopsinae. Tulane Stiidies in Geology and Paleontology 26: 49-160.
Vokes E.H. 1995. Two new Cenozoic Muricinae (Gastropoda: Murieidae) of the western Atlantic région. Tulane Stiidies in Geology and Paleontology 28: 119-122.
35
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C. VlLVBNS
Novapex 15(2): 37-48, K) juin 2014
New species and new records of Calliostomatidae (Gastropoda: Trochoidea) from eastern and central Indo-Pacific
Claude VILVENS
Rue de Hermalle, 1 13 - B-4680 Oupeye, Belgium Scientific Collaborator, Muséum national d'Histoire naturelle, Paris.
vilvens.claude(g>skynet.be
KEYWORDS. Gastropoda, Calliostomatidae, Tonga Islands, Austral Archipelago, Vanuatu, Solomon Islands, Calliostoma, new species.
ABSTRACT. New records of five known Calliostomatidae species from eastern and central tropical Pacific are listed, extending the distribution area of some of them. Four new species are described and compared with similar species; Calliostoma haapaiensis n. sp., C. vaubanoides n. sp., C. mesemorinon n. sp. and C . polysarkon n. sp.
RESUME. De nouveaux relevés de cinq espèces connues de Calliostomatidae provenant de l'est et du centre du Pacifique tropical sont listés, étendant ainsi faire de distribution d'un certain nombre d'entre elles. Quatre nouvelles espèces sont décrites et comparées avec des espèces similaires : Calliostoma haapaiensis n. sp., C. vaubanoides n. sp., C. mesemorinon n. sp. et C. polysarkon n. sp.
INTRODUCTION
The malacofauna, especially Trochoidea species, of eastern tropical Pacific is still rather poorly known. Only a few dredging, like those of IRD (Institut de Recherche pour le Développement, Paris - ex- ORSTOM) and MNHN (Muséum national d'Histoire naturelle, Paris) campaigns in French Polynesia (Vilvens, 2012) or in Tonga Islands (Vilvens, 2005), and local samplings (Stratmann & Stahlschmidt, 2007, Vilvens, 2009a) hâve brought a limited amount of samples.
On the contrary, numerous campaigns hâve been conducted in central tropical Pacific, especially again the ones organized by IRD-MNHN in New Caledonia, Vanuatu, Fiji and Solomon Islands, bringing a huge material (Héros et al., 2007; Bouchet et al., 2008), notably Calliostomatidae (Trochoidea) samples with numerous new species (e.g.: Marshall, 1995; Vilvens, 2009b).
While studying varions Trochoidea and Seguenzoidea from ail these campaigns, a few Calliostomatidae samples were found among other
trochids samples, bringing additional records of known species but also more surprisingly some new species. The présent paper présents the results of this recent study.
Material and methods
The material studied in the présent paper was brought mainly by some IRD-MNHN expéditions: BORDAU
2 (6/2000), BENTHAUS (11/2002), SALOMON 2 (10-11/2004), BOA 1 (9/2005) and SALOMONBOA
3 (9-10/2007) (see Map 1).
Regarding the distribution of the new species and the extension of the distribution of known species, the range is taken from the internai intervals of the two extremes values. This range of the known and new species is provided for ail the available specimens and also for the only living specimens if they hâve been found; when these ranges are the same, the common range is cited once with the "(living)" annotation; if ail the specimens are dead collected, the range is cited with the "(dead)" annotation.
37
C. VlLVENS
New species and new records of Cailiostomatidae
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Map 1. Records of cited Cailiostomatidae species in eastern and central tropical Pacific :
★ Calliostoma mesemorinon n. sp. (BENTHAUS)
® C . paradigmaîum Marshall, 1995 (BENTHAUS)
* C. arx Vilvens, 2005 (BORDAU 2)
+ C. haapaiensis n. sp. (BORDAU 2)
• C. vaubanoides n. sp. (BORDAU 2)
♦ C. hexalyssion Vilvens, 2009 (SALOMON 2)
□ C. aporia Vilvens, 2009 (SALOMONBOA 3)
O C. siiduirauti Bozzetti, 1997 (SALOMONBOA 3) 0 C . polysarkon n. sp. (BOA 1)
Regarding the description methodology, the main conchological features used are (see Eig. 1 below):
♦ general shape of the shell (spire high spired or depressed - conical, cyrtoconoidal, coeloconoidal);
♦ size and shape of the protoconch;
♦ shape of the whorls (convex, concave, straight - with or without shoulder or keel);
♦ spiral cords of the whorls (ontogeny, number, beaded or smooth, distance between cords);
♦ shape of the aperture, features of the outer and the inner lip;
♦ shape of the base and spiral cords : number, beaded or smooth, distance between cords;
♦ features of the umbilicus : open or covered with a callus, relative size, spiral cords around or inside LimbilicLis;
♦ columella ; thickened or not;
♦ colour of the protoconch, of the whorls, of the base.
38
C. ViLVENS
Novapex 15(2): 37-48, 10 juin 2014
Figure 1, Features of Calliostomatidae shells; FI : height; W ; width; HA : height of the aperture; PI , P2, P3, ... : primary cords; SI, S2, S3, ... : secondary cords; Tl, T2, ... : tertiaiy cords (shell : Calliostoma (Fautor) chloriim Vilvens, 2005, Fiji, BORD AU l,stnDW1454, 13.6 x 10.4 mm).
Abbreviations
Repositories
IMT: Institute of Malacology of Tokyo, Tokyo, Japan. MNHN: Muséum national d'Histoire naturelle, Paris, France.
Other abbreviations (see Fig. 1 above)
H: height.
W: width.
HA; height of the aperture.
TW; number of teleoconeh whorls.
PI , P2, P3, ...: primary cords (PI is the most adapical). Pi: generic name for the primary cords (i=l,2,3, ...). SI, S2, S3, ...: secondary cords (SI is the most adapical).
Si: generic name for the seeondary cords (i=l ,2,3, ...). Tl, T2, T3, ...: tertiary cords (numbered following appearing order).
stn: station.
Iv: live-taken specimens présent in sample. dd: no live-taken specimens présent in sample. sub: subadult specimen. juv: juvénile specimen.
SYSTEMATICS
We follow bere Marshall (1995), Bouchet & Roeroi (2005) and Williams et al. (2008, 2010) where Calliostomatidae, earlier treated as a subfamily of Trochidae (Hiekman & MeLean, 1990), are now ranked as a family of superfamily Troehoidea, with the two SLibfamilies Calliostomatinae and Thysanodontinae .
Regarding subgenera of the genus Calliostoma , we décidé to not use them here, beeause they seem today rather artificial considering the information brought by DNA studies. One ean however refer to Marshall (1995) regarding Indo-Pacifie subgenera.
39
C. ViLVENS
New species and new records of Calliostomatidae
Superfamily TROCHOIDEA Rafinesque, 1815 Family CALLIOSTOMATIDAE Thiele, 1924 Subfamily CALLIOSTOMATINAE Thiele, 1924 Tribe Calliostomatini Thiele, 1924 [= Ziziphininae Gray, 1847 J
Genus Calliostoma Swainson, 1840
Type species Trochus coniiliis Linnaeus, 1758 (by s.d.
Herrmannsen, 1846) - Recent, Mediteiranean Sea.
Calliostoma arx Vilvens, 2005 Figs 30-31
Calliostoma (Benthastelena) arx Vilvens, 2005: 14- 16, figs 41-44. Type locality: Tonga Islands, south of Eua Island, 483-531 m.
Material examined. Tonga Islands. BORDAU 2: stn CP1644,21°05'S, 175°23'W,501 m, 1 dd.
Distribution. Tonga, 483-531 m (dead); Fiji, 450- 500 m (dead).
Remarks. The main characteristics of this species are:
- height Lip to 18 mm, width up to 16 mm;
- coeloconidal in shape, slightly higher than wide; angulate periphery;
- protoconch about 350 /im wide, of 1 .25 whorl;
- teleoconch with up to 8.5 convex whorls with spiny spiral cords; P2 and P3 appearing first while PI appears a quarter of whorl later; P4 slightly emerging from suture on fourth whorl; S4 and SI appearing on sixth whorl, S2 on seventh whorl, S3 absent;
- weakly convex base with 12 rather thin granular spiral cords;
- narrow and deep umbilicus;
- cream white.
Calliostoma haapaiensis n. sp.
Figs 2-4
Type material. Holotype (14.1 x 1 1 .5 mm) MNHN (lM-2000-27238).
Type locality. Tonga Islands, north of Ha'apai group, BORDAU 2, stn DW1595, 19°03'S, 174°19'W, 523- 806 m.
Distribution. Tonga Islands, 523-806 m (dead).
Diagnosis. A Calliostoma species of medium size with a conical spire, concave whorls with up to 14 granular, close spiral cords, the two most abapical cords being the strongest and making a keel, a weakly convex base with up to 20 granular spiral cords, an umbilicus partly closed by the columella, an off white colour with irregular brownish orange liâmes and a peripheral cord alternating white and brown segments.
Description. Shell of medium size for the genus (height up to 14.1 mm, width up to 1 1 .5 mm), higher
than wide, conical to weakly coeloconoidal in shape; spire elevated, height 1 .2x width, 4.1x aperture height; angulate periphery; umbilicate.
Protoconch about 300 //m wide, of 1.25 whorl, roLinded, covered by a network of ridges producing large polygonal areas; thin, poorly visible terminal varix.
Teleoconch of 8.5 whorls, early whorls fiat, next ones concave. Suture very poorly visible, not canaliculate. First whorl moderately convex, sculptured by rather thick, almost orthocline ribs and 3 spiral cords; P2 and P3 appearing immediately, PI appearing a quarter of whorl later; P3 stronger and PI weaker than other cords; distance between cords about 3x width of cords; distance between ribs 4x width of ribs; axial ribs making the cords granular. Second whorl only weakly convex to almost fiat; ail cords stronger, with thick beads; P3 very stronger than other cords, with pointed beads; distance between PI and P2 similar in size to cords, distance between P2 and P3 about 2x width of P2; axial ribs prosocline; suture poorly visible. Third whorl fiat with P3 much stronger with Sharp beads, producing keel; PI weaker than P3, but stronger than P2; S2 appearing, very thin; P4 partly visible, almost completely hidden by succeeding whorl; axial ribs still strong, distance between similar in size to width of ribs. On fourth whorl, SI appearing; S2 similar in size to P2; Tl appearing between P2 and S2; P4 almost fully visible; beads of P3 thick, bluntly pointed; axial ribs between P2 and P3 thicker than other cords; whorl slightly concave in shape, with a basal keel made by P3. On fifth whorl, beads of PI axially elongated; ail cords similar in size except PI stronger and P3 much stronger; axial ribs weaker, reduced to thin threads; whorl strongly concave between suture and P3; P3 and P4 making a strong peripheral keel. On sixth whorl, T2 appearing between S2 and P3, quickly as strong as S2; axial threads no more visible. On last whorls, additional thin, smooth cords appearing elsewhere between PI and P3, number of cords reaching about 14; angular periphery.
Aperture subquadrangular; outer lip thin, curved, with a basal part rounded, meeting inner lip with rounded angle. Columella oblique, almost straight, without tooth.
Base weakly convex, with about 20 spiral cords; outer cords very thin, subgranular; cords thickening towards umbilicus and becoming granular, innermost cords twice thicker than outermost cords; fine axial threads on the whole surface.
Umbilicus narrow (about 7% of shell width), funnel shaped with gentle slope, half covered by an expansion of columella.
Colour of teleoconch global ly off white, with brownish orange fiâmes; P3 with regular brown patches covering two or three beads and separated by four or five whitish coloured beads; base nacreous pinkish white; protoconch white translucent.
40
C. ViLVENS
Novapex 15(2): 37-48, 10 juin 2014
Discussion. Calliostoma haapaiensis n. sp. is rallier close to C. héros Marshall, 1995 from Loyalty Islands (Figs 5-7), but this similar in size and in spiral cords ontogeny species lias a smaller H/W ratio for specimens of the same size, a protoconch with a strong terminal varix, only weakly (not strongly) concave vvhorls with a very much less prominent peripheral keel, thicker spiral cords on whorls and base, thicker and not similar in size beads on P3.
The new species is close to C. katoi Sakurai, 1994 from Japan (Figs 8-9) but this similar in size species lias an umbilicus, has less numerous, thicker spiral cords on the whorls and only 6-7 sniooth spiral cords on the base.
Etymology. From the type locality, Ha'apai islands, one of the three islands groups of Tonga archipelago.
Calliostoirm vaubanoides n. sp.
Figs 14-16
Type materiai. Holotype (14.8 x 15.3 mm) MNHN (lM-2000-27239).
Type locality. Tonga Islands, south of Nomuka group, BORDAU 2, stn DW1548, 20°38'S, 175°03'W, 476- 478 m.
Distribution. Tonga Islands, 476-478 m (dead).
Diagnosis. A Calliostoma species of medium size with a coeloconoidal spire, fiat whorls with up to 9 granular spiral cords, the peripheral cords being the strongest and making a weak keel, an almost fiat base with up to 14 granular spiral cords, no umbilicus, an orange colour with brow fiâmes on the adapical area of the whorls and on the base.
Description, Shell of medium size for the genus (height up to 14.8 mm, width up to 15.3 mm), wider than high, coeloconoidal in shape; spire moderately elevated, height 0.97x width, 3.0x aperture height; angulate periphery; anomphalous.
Protoconch about 350 pm wide, of 1.25 whorl, rounded, covered by a network of ridges producing large polygonal areas; thin terminal varix.
Teleoconch of 7.0 whorls, ail almost fiat except the two last whorls weakly convex.
Suture visible, not canaliculate.
First whorl moderately convex, sculptured by prosocline ribs and 3 spiral cords; P2 and P3 appearing immediately, PI appearing about a quarter of whorl later; P4 completely hidden by succeeding whorl; PI slightly weaker than other cords; distance between cords about 2x width of cords; distance between ribs 1 .5x width of ribs; axial ribs making the cords granular. Second whorl only weakly convex to almost fiat; ail cords stronger and similar in size, with thick beads; axial ribs similar in thickness to the cords, giving a reticulate pattern. Third whorl fiat in shape;
P2 weaker than other cords, P3 the strongest with weakly pointed beads; S2 appearing at mid whorl, very thin; S 1 appearing at end of whorl, also very thin. On foLirth whorl, S3 appearing; SI and S2 still much thinner than Pi; PI and P3 stronger than ail other cords, P3 the strongest with bluntly pointed beads, making a weak keel; axial threads weakening. On fifth whorl, axial threads disappearing; ail cords similar in size except PI stronger and P3 the strongest. On sixth whorl, Tl appearing between PI and SI and T2 between SI and P2; P4 partially emerging from suture; beads of ail the cords pointed. On last whorl, S2, P3 and S 3 stronger than other cords; P4 very weaker than other cords; angular periphery.
Aperture subtriangular; outer lip rather thin, slightly curved, with a basal part almost straight, meeting inner lip with a distinct angle. Columella vertical, slightly curved, without tooth.
Base very weakly convex to almost fiat, with 14 spiral cords; innermost cords slightly thicker than outermost, ail granular; interspaces between cords as wide as cords. No umbilicus.
Colour of two first teleoconch whorls off white, other whorls orange with brow fiâmes on the adapical area; base off white to light orange, with axial brown fiâmes; protoconch off white.
Discussion. Calliostoma vaubanoides n. sp. is close to Calliostoma vaubani Marshall, 1995 from Northern New Caledonia (Figs 10-11), but the latter species is smaller for a similar number of whorls, is higher than wide, has a larger protoconch (400-430 //m) and has a different spiral cords ontogeny with SI appearing much more later,
The new species looks also a little like Calliostoma monikae Stratmann & Schwabe, 2007 from Samoa Islands, but the latter species is smaller (height up to 8.5 mm) for a similar number of whorls, is higher than wide with a strictly conical shape and an oblique, thickened columella.
Etymology. Similar in shape (Greek suffix: -oiôqQ - from the closest Calliostoma species, C. vaubani Marshall, 1995.
Calliostoma mesemorinon n. sp.
Figs 17-19, Table 1
Type materiai. Holotype (13.3 x 10.6 mm) MNHN (lM-2000-27240). Paratypes 2 MNHN (IM-2000- 27241).
Type locality. French Polynesia, Austral Archipelago, eastern coast of Rurutu, BENTHAUS, stn CAS2008, 22°27'S, 1 5 ri9'W, 280-300 m.
Materiai examined. Austral Archipelago.
BENTHAUS; stn CAS2008, 22°27'S, 151°19'W, 280- 300 m, 4 dd (holotype and 2 paratypes).
41
C. VlLVENS
New species and new records of Calliostomatidae
Distribution. French Polynesia, Austral Archipelago, 280-300 m (dead).
Diagnosis. A Calliostoma species of medium size, with a conical spire, up to 5 strongly granular, close spiral cords, the most adapical cord finally the strongest, an almost fiat base with up to